Publications

Displaying 301 - 400 of 812
  • Hubers, F., Trompenaars, T., Collin, S., De Schepper, K., & De hoop, H. (2020). Hypercorrection as a by-product of education. Applied Linguistics, 41(4), 552-574. doi:10.1093/applin/amz001.

    Abstract

    Prescriptive grammar rules are taught in education, generally to ban the use of certain frequently encountered constructions in everyday language. This may lead to hypercorrection, meaning that the prescribed form in one construction is extended to another one in which it is in fact prohibited by prescriptive grammar. We discuss two such cases in Dutch: the hypercorrect use of the comparative particle dan ‘than’ in equative constructions, and the hypercorrect use of the accusative pronoun hen ‘them’ for a dative object. In two experiments, high school students of three educational levels were tested on their use of these hypercorrect forms (nexp1 = 162, nexp2 = 159). Our results indicate an overall large amount of hypercorrection across all levels of education, including pre-university level students who otherwise perform better in constructions targeted by prescriptive grammar rules. We conclude that while teaching prescriptive grammar rules to high school students seems to increase their use of correct forms in certain constructions, this comes at a cost of hypercorrection in others.
  • Hubers, F. (2020). Two of a kind: Idiomatic expressions by native speakers and second language learners. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Huettig, F., Guerra, E., & Helo, A. (2020). Towards understanding the task dependency of embodied language processing: The influence of colour during language-vision interactions. Journal of Cognition, 3(1): 41. doi:10.5334/joc.135.

    Abstract

    A main challenge for theories of embodied cognition is to understand the task dependency of embodied language processing. One possibility is that perceptual representations (e.g., typical colour of objects mentioned in spoken sentences) are not activated routinely but the influence of perceptual representation emerges only when context strongly supports their involvement in language. To explore this question, we tested the effects of colour representations during language processing in three visual- world eye-tracking experiments. On critical trials, participants listened to sentence- embedded words associated with a prototypical colour (e.g., ‘...spinach...’) while they inspected a visual display with four printed words (Experiment 1), coloured or greyscale line drawings (Experiment 2) and a ‘blank screen’ after a preview of coloured or greyscale line drawings (Experiment 3). Visual context always presented a word/object (e.g., frog) associated with the same prototypical colour (e.g. green) as the spoken target word and three distractors. When hearing spinach participants did not prefer the written word frog compared to other distractor words (Experiment 1). In Experiment 2, colour competitors attracted more overt attention compared to average distractors, but only for the coloured condition and not for greyscale trials. Finally, when the display was removed at the onset of the sentence, and in contrast to the previous blank-screen experiments with semantic competitors, there was no evidence of colour competition in the eye-tracking record (Experiment 3). These results fit best with the notion that the main role of perceptual representations in language processing is to contextualize language in the immediate environment.

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    Data files and script
  • Huizeling, E., Wang, H., Holland, C., & Kessler, K. (2020). Age-related changes in attentional refocusing during simulated driving. Brain sciences, 10(8): 530. doi:10.3390/brainsci10080530.

    Abstract

    We recently reported that refocusing attention between temporal and spatial tasks becomes more difficult with increasing age, which could impair daily activities such as driving (Callaghan et al., 2017). Here, we investigated the extent to which difficulties in refocusing attention extend to naturalistic settings such as simulated driving. A total of 118 participants in five age groups (18–30; 40–49; 50–59; 60–69; 70–91 years) were compared during continuous simulated driving, where they repeatedly switched from braking due to traffic ahead (a spatially focal yet temporally complex task) to reading a motorway road sign (a spatially more distributed task). Sequential-Task (switching) performance was compared to Single-Task performance (road sign only) to calculate age-related switch-costs. Electroencephalography was recorded in 34 participants (17 in the 18–30 and 17 in the 60+ years groups) to explore age-related changes in the neural oscillatory signatures of refocusing attention while driving. We indeed observed age-related impairments in attentional refocusing, evidenced by increased switch-costs in response times and by deficient modulation of theta and alpha frequencies. Our findings highlight virtual reality (VR) and Neuro-VR as important methodologies for future psychological and gerontological research.

    Additional information

    supplementary file
  • Iacozza, S., Meyer, A. S., & Lev-Ari, S. (2020). How in-group bias influences the level of detail of speaker-specific information encoded in novel lexical representations. Journal of Experimental Psychology: Learning, Memory, and Cognition, 46(5), 894-906. doi:10.1037/xlm0000765.

    Abstract

    An important issue in theories of word learning is how abstract or context-specific representations of novel words are. One aspect of this broad issue is how well learners maintain information about the source of novel words. We investigated whether listeners’ source memory was better for words learned from members of their in-group (students of their own university) than it is for words learned from members of an out-group (students from another institution). In the first session, participants saw 6 faces and learned which of the depicted students attended either their own or a different university. In the second session, they learned competing labels (e.g., citrus-peller and citrus-schiller; in English, lemon peeler and lemon stripper) for novel gadgets, produced by the in-group and out-group speakers. Participants were then tested for source memory of these labels and for the strength of their in-group bias, that is, for how much they preferentially process in-group over out-group information. Analyses of source memory accuracy demonstrated an interaction between speaker group membership status and participants’ in-group bias: Stronger in-group bias was associated with less accurate source memory for out-group labels than in-group labels. These results add to the growing body of evidence on the importance of social variables for adult word learning.
  • Iacozza, S. (2020). Exploring social biases in language processing. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Indefrey, P., & Levelt, W. J. M. (1999). A meta-analysis of neuroimaging experiments on word production. Neuroimage, 7, 1028.
  • Indefrey, P. (1998). De neurale architectuur van taal: Welke hersengebieden zijn betrokken bij het spreken. Neuropraxis, 2(6), 230-237.
  • Indefrey, P., Gruber, O., Brown, C. M., Hagoort, P., Posse, S., & Kleinschmidt, A. (1998). Lexicality and not syllable frequency determine lateralized premotor activation during the pronunciation of word-like stimuli: An fMRI study. NeuroImage, 7, S4.
  • Indefrey, P. (1999). Some problems with the lexical status of nondefault inflection. Behavioral and Brain Sciences, 22(6), 1025. doi:10.1017/S0140525X99342229.

    Abstract

    Clahsen's characterization of nondefault inflection as based exclusively on lexical entries does not capture the full range of empirical data on German inflection. In the verb system differential effects of lexical frequency seem to be input-related rather than affecting morphological production. In the noun system, the generalization properties of -n and -e plurals exceed mere analogy-based productivity.
  • Isbilen, E. S., McCauley, S. M., Kidd, E., & Christiansen, M. H. (2020). Statistically induced chunking recall: A memory‐based approach to statistical learning. Cognitive Science, 44(7): e12848. doi:10.1111/cogs.12848.

    Abstract

    The computations involved in statistical learning have long been debated. Here, we build on work suggesting that a basic memory process, chunking , may account for the processing of statistical regularities into larger units. Drawing on methods from the memory literature, we developed a novel paradigm to test statistical learning by leveraging a robust phenomenon observed in serial recall tasks: that short‐term memory is fundamentally shaped by long‐term distributional learning. In the statistically induced chunking recall (SICR) task, participants are exposed to an artificial language, using a standard statistical learning exposure phase. Afterward, they recall strings of syllables that either follow the statistics of the artificial language or comprise the same syllables presented in a random order. We hypothesized that if individuals had chunked the artificial language into word‐like units, then the statistically structured items would be more accurately recalled relative to the random controls. Our results demonstrate that SICR effectively captures learning in both the auditory and visual modalities, with participants displaying significantly improved recall of the statistically structured items, and even recall specific trigram chunks from the input. SICR also exhibits greater test–retest reliability in the auditory modality and sensitivity to individual differences in both modalities than the standard two‐alternative forced‐choice task. These results thereby provide key empirical support to the chunking account of statistical learning and contribute a valuable new tool to the literature.
  • Jacoby, N., Margulis, E. H., Clayton, M., Hannon, E., Honing, H., Iversen, J., Klein, T. R., Mehr, S. A., Pearson, L., Peretz, I., Perlman, M., Polak, R., Ravignani, A., Savage, P. E., Steingo, G., Stevens, C. J., Trainor, L., Trehub, S., Veal, M., & Wald-Fuhrmann, M. (2020). Cross-cultural work in music cognition: Challenges, insights, and recommendations. Music Perception, 37(3), 185-195. doi:10.1525/mp.2020.37.3.185.

    Abstract

    Many foundational questions in the psychology of music require cross-cultural approaches, yet the vast majority of work in the field to date has been conducted with Western participants and Western music. For cross-cultural research to thrive, it will require collaboration between people from different disciplinary backgrounds, as well as strategies for overcoming differences in assumptions, methods, and terminology. This position paper surveys the current state of the field and offers a number of concrete recommendations focused on issues involving ethics, empirical methods, and definitions of “music” and “culture.”
  • Janse, E., & Quené, H. (1999). On the suitability of the cross-modal semantic priming task. In Proceedings of the XIVth International Congress of Phonetic Sciences (pp. 1937-1940).
  • Janssen, D. (1999). Producing past and plural inflections. PhD Thesis, Radboud University Nijmegen, Nijmegen. doi:10.17617/2.2057667.
  • Jebb, D., Huang, Z., Pippel, M., Hughes, G. M., Lavrichenko, K., Devanna, P., Winkler, S., Jermiin, L. S., Skirmuntt, E. C., Katzourakis, A., Burkitt-Gray, L., Ray, D. A., Sullivan, K. A. M., Roscito, J. G., Kirilenko, B. M., Dávalos, L. M., Corthals, A. P., Power, M. L., Jones, G., Ransome, R. D. and 9 moreJebb, D., Huang, Z., Pippel, M., Hughes, G. M., Lavrichenko, K., Devanna, P., Winkler, S., Jermiin, L. S., Skirmuntt, E. C., Katzourakis, A., Burkitt-Gray, L., Ray, D. A., Sullivan, K. A. M., Roscito, J. G., Kirilenko, B. M., Dávalos, L. M., Corthals, A. P., Power, M. L., Jones, G., Ransome, R. D., Dechmann, D., Locatelli, A. G., Puechmaille, S. J., Fedrigo, O., Jarvis, E. D., Hiller, M., Vernes, S. C., Myers, E. W., & Teeling, E. C. (2020). Six reference-quality genomes reveal evolution of bat adaptations. Nature, 583, 578-584. doi:10.1038/s41586-020-2486-3.

    Abstract

    Bats possess extraordinary adaptations, including flight, echolocation, extreme longevity and unique immunity. High-quality genomes are crucial for understanding the molecular basis and evolution of these traits. Here we incorporated long-read sequencing and state-of-the-art scaffolding protocols1 to generate, to our knowledge, the first reference-quality genomes of six bat species (Rhinolophus ferrumequinum, Rousettus aegyptiacus, Phyllostomus discolor, Myotis myotis, Pipistrellus kuhlii and Molossus molossus). We integrated gene projections from our ‘Tool to infer Orthologs from Genome Alignments’ (TOGA) software with de novo and homology gene predictions as well as short- and long-read transcriptomics to generate highly complete gene annotations. To resolve the phylogenetic position of bats within Laurasiatheria, we applied several phylogenetic methods to comprehensive sets of orthologous protein-coding and noncoding regions of the genome, and identified a basal origin for bats within Scrotifera. Our genome-wide screens revealed positive selection on hearing-related genes in the ancestral branch of bats, which is indicative of laryngeal echolocation being an ancestral trait in this clade. We found selection and loss of immunity-related genes (including pro-inflammatory NF-κB regulators) and expansions of anti-viral APOBEC3 genes, which highlights molecular mechanisms that may contribute to the exceptional immunity of bats. Genomic integrations of diverse viruses provide a genomic record of historical tolerance to viral infection in bats. Finally, we found and experimentally validated bat-specific variation in microRNAs, which may regulate bat-specific gene-expression programs. Our reference-quality bat genomes provide the resources required to uncover and validate the genomic basis of adaptations of bats, and stimulate new avenues of research that are directly relevant to human health and disease

    Additional information

    41586_2020_2486_MOESM1_ESM.pdf
  • Jessop, A., & Chang, F. (2020). Thematic role information is maintained in the visual object-tracking system. Quarterly journal of experimental psychology, 73(1), 146-163. doi:10.1177%2F1747021819882842.

    Abstract

    Thematic roles characterise the functions of participants in events, but there is no agreement on how these roles are identified in the real world. In three experiments, we examined how role identification in push events is supported by the visual object-tracking system. Participants saw one to three push events in visual scenes with nine identical randomly moving circles. After a period of random movement, two circles from one of the push events and a foil object were given different colours and the participants had to identify their roles in the push with an active sentence, such as red pushed blue. It was found that the participants could track the agent and patient targets and generate descriptions that identified their roles at above chance levels, even under difficult conditions, such as when tracking multiple push events (Experiments 1–3), fixating their gaze (Experiment 1), performing a concurrent speeded-response task (Experiment 2), and when tracking objects that were temporarily invisible (Experiment 3). The results were consistent with previous findings of an average tracking capacity limit of four objects, individual differences in this capacity, and the use of attentional strategies. The studies demonstrated that thematic role information can be maintained when tracking the identity of visually identical objects, then used to map role fillers (e.g., the agent of a push event) into their appropriate sentence positions. This suggests that thematic role features are stored temporarily in the visual object-tracking system.
  • Jongman, S. R., Roelofs, A., & Lewis, A. G. (2020). Attention for speaking: Prestimulus motor-cortical alpha power predicts picture naming latencies. Journal of Cognitive Neuroscience, 32(5), 747-761. doi:10.1162/jocn_a_01513.

    Abstract

    There is a range of variability in the speed with which a single speaker will produce the same word from one instance to another. Individual differences studies have shown that the speed of production and the ability to maintain attention are related. This study investigated whether fluctuations in production latencies can be explained by spontaneous fluctuations in speakers' attention just prior to initiating speech planning. A relationship between individuals' incidental attentional state and response performance is well attested in visual perception, with lower prestimulus alpha power associated with faster manual responses. Alpha is thought to have an inhibitory function: Low alpha power suggests less inhibition of a specific brain region, whereas high alpha power suggests more inhibition. Does the same relationship hold for cognitively demanding tasks such as word production? In this study, participants named pictures while EEG was recorded, with alpha power taken to index an individual's momentary attentional state. Participants' level of alpha power just prior to picture presentation and just prior to speech onset predicted subsequent naming latencies. Specifically, higher alpha power in the motor system resulted in faster speech initiation. Our results suggest that one index of a lapse of attention during speaking is reduced inhibition of motor-cortical regions: Decreased motor-cortical alpha power indicates reduced inhibition of this area while early stages of production planning unfold, which leads to increased interference from motor-cortical signals and longer naming latencies. This study shows that the language production system is not impermeable to the influence of attention.
  • Jongman, S. R., Piai, V., & Meyer, A. S. (2020). Planning for language production: The electrophysiological signature of attention to the cue to speak. Language, Cognition and Neuroscience, 35(7), 915-932. doi:10.1080/23273798.2019.1690153.

    Abstract

    In conversation, speech planning can overlap with listening to the interlocutor. It has been
    postulated that once there is enough information to formulate a response, planning is initiated
    and the response is maintained in working memory. Concurrently, the auditory input is
    monitored for the turn end such that responses can be launched promptly. In three EEG
    experiments, we aimed to identify the neural signature of phonological planning and monitoring
    by comparing delayed responding to not responding (reading aloud, repetition and lexical
    decision). These comparisons consistently resulted in a sustained positivity and beta power
    reduction over posterior regions. We argue that these effects reflect attention to the sequence
    end. Phonological planning and maintenance were not detected in the neural signature even
    though it is highly likely these were taking place. This suggests that EEG must be used cautiously
    to identify response planning when the neural signal is overridden by attention effects
  • Jordanoska, I. (2020). The pragmatics of sentence final and second position particles in Wolof. PhD Thesis, University of Vienna, Vienna.
  • Jordens, P. (1998). Defaultformen des Präteritums. Zum Erwerb der Vergangenheitsmorphologie im Niederlänidischen. In H. Wegener (Ed.), Eine zweite Sprache lernen (pp. 61-88). Tübingen, Germany: Verlag Gunter Narr.
  • Kastens, K. (2020). The Jerome Bruner Library treasure. In M. E. Poulsen (Ed.), The Jerome Bruner Library: From New York to Nijmegen (pp. 29-34). Nijmegen: Max Planck Institute for Psycholinguistics.
  • Kaufeld, G., Naumann, W., Meyer, A. S., Bosker, H. R., & Martin, A. E. (2020). Contextual speech rate influences morphosyntactic prediction and integration. Language, Cognition and Neuroscience, 35(7), 933-948. doi:10.1080/23273798.2019.1701691.

    Abstract

    Understanding spoken language requires the integration and weighting of multiple cues, and may call on cue integration mechanisms that have been studied in other areas of perception. In the current study, we used eye-tracking (visual-world paradigm) to examine how contextual speech rate (a lower-level, perceptual cue) and morphosyntactic knowledge (a higher-level, linguistic cue) are iteratively combined and integrated. Results indicate that participants used contextual rate information immediately, which we interpret as evidence of perceptual inference and the generation of predictions about upcoming morphosyntactic information. Additionally, we observed that early rate effects remained active in the presence of later conflicting lexical information. This result demonstrates that (1) contextual speech rate functions as a cue to morphosyntactic inferences, even in the presence of subsequent disambiguating information; and (2) listeners iteratively use multiple sources of information to draw inferences and generate predictions during speech comprehension. We discuss the implication of these demonstrations for theories of language processing
  • Kaufeld, G., Ravenschlag, A., Meyer, A. S., Martin, A. E., & Bosker, H. R. (2020). Knowledge-based and signal-based cues are weighted flexibly during spoken language comprehension. Journal of Experimental Psychology: Learning, Memory, and Cognition, 46(3), 549-562. doi:10.1037/xlm0000744.

    Abstract

    During spoken language comprehension, listeners make use of both knowledge-based and signal-based sources of information, but little is known about how cues from these distinct levels of representational hierarchy are weighted and integrated online. In an eye-tracking experiment using the visual world paradigm, we investigated the flexible weighting and integration of morphosyntactic gender marking (a knowledge-based cue) and contextual speech rate (a signal-based cue). We observed that participants used the morphosyntactic cue immediately to make predictions about upcoming referents, even in the presence of uncertainty about the cue’s reliability. Moreover, we found speech rate normalization effects in participants’ gaze patterns even in the presence of preceding morphosyntactic information. These results demonstrate that cues are weighted and integrated flexibly online, rather than adhering to a strict hierarchy. We further found rate normalization effects in the looking behavior of participants who showed a strong behavioral preference for the morphosyntactic gender cue. This indicates that rate normalization effects are robust and potentially automatic. We discuss these results in light of theories of cue integration and the two-stage model of acoustic context effects
  • Kaufeld, G., Bosker, H. R., Ten Oever, S., Alday, P. M., Meyer, A. S., & Martin, A. E. (2020). Linguistic structure and meaning organize neural oscillations into a content-specific hierarchy. The Journal of Neuroscience, 49(2), 9467-9475. doi:10.1523/JNEUROSCI.0302-20.2020.

    Abstract

    Neural oscillations track linguistic information during speech comprehension (e.g., Ding et al., 2016; Keitel et al., 2018), and are known to be modulated by acoustic landmarks and speech intelligibility (e.g., Doelling et al., 2014; Zoefel & VanRullen, 2015). However, studies investigating linguistic tracking have either relied on non-naturalistic isochronous stimuli or failed to fully control for prosody. Therefore, it is still unclear whether low frequency activity tracks linguistic structure during natural speech, where linguistic structure does not follow such a palpable temporal pattern. Here, we measured electroencephalography (EEG) and manipulated the presence of semantic and syntactic information apart from the timescale of their occurrence, while carefully controlling for the acoustic-prosodic and lexical-semantic information in the signal. EEG was recorded while 29 adult native speakers (22 women, 7 men) listened to naturally-spoken Dutch sentences, jabberwocky controls with morphemes and sentential prosody, word lists with lexical content but no phrase structure, and backwards acoustically-matched controls. Mutual information (MI) analysis revealed sensitivity to linguistic content: MI was highest for sentences at the phrasal (0.8-1.1 Hz) and lexical timescale (1.9-2.8 Hz), suggesting that the delta-band is modulated by lexically-driven combinatorial processing beyond prosody, and that linguistic content (i.e., structure and meaning) organizes neural oscillations beyond the timescale and rhythmicity of the stimulus. This pattern is consistent with neurophysiologically inspired models of language comprehension (Martin, 2016, 2020; Martin & Doumas, 2017) where oscillations encode endogenously generated linguistic content over and above exogenous or stimulus-driven timing and rhythm information.
  • Keating, E. (1995). Pilot questionnaire to investigate social uses of space, especially as related to 1) linguistic practices and 2) social organization. In D. Wilkins (Ed.), Extensions of space and beyond: manual for field elicitation for the 1995 field season (pp. 17-21). Nijmegen: Max Planck Institute for Psycholinguistics. doi:10.17617/2.3004227.

    Abstract

    Day-to-day interpretations of “space” are enmeshed in specific cultural and linguistic practices. For example, many cultures have an association between vertical height and social standing; more powerful people may be placed literally higher than others at social gatherings, and be spoken of as having higher status. This questionnaire is a guide for exploring relationships between space, language, and social structure. The goal is to better understand how space is organised in the focus community, and to investigate the extent to which space is used as a model for reproducing social forms.
  • Kempen, G. (1991). Conjunction reduction and gapping in clause-level coordination: An inheritance-based approach. Computational Intelligence, 7, 357-360. doi:10.1111/j.1467-8640.1991.tb00406.x.
  • Kempen, G. (1995). De mythe van het woordbeeld: Spellingherziening taalpsychologisch doorgelicht. Onze Taal, 64(11), 275-277.
  • Kempen, G. (1995). Drinken eten mij Nim. Intermediair, 31(19), 41-45.
  • Kempen, G. (1998). Comparing and explaining the trajectories of first and second language acquisition: In search of the right mix of psychological and linguistic factors [Commentory]. Bilingualism: Language and Cognition, 1, 29-30. doi:10.1017/S1366728998000066.

    Abstract

    When you compare the behavior of two different age groups which are trying to master the same sensori-motor or cognitive skill, you are likely to discover varying learning routes: different stages, different intervals between stages, or even different orderings of stages. Such heterogeneous learning trajectories may be caused by at least six different types of factors: (1) Initial state: the kinds and levels of skills the learners have available at the onset of the learning episode. (2) Learning mechanisms: rule-based, inductive, connectionist, parameter setting, and so on. (3) Input and feedback characteristics: learning stimuli, information about success and failure. (4) Information processing mechanisms: capacity limitations, attentional biases, response preferences. (5) Energetic variables: motivation, emotional reactions. (6) Final state: the fine-structure of kinds and levels of subskills at the end of the learning episode. This applies to language acquisition as well. First and second language learners probably differ on all six factors. Nevertheless, the debate between advocates and opponents of the Fundamental Difference Hypothesis concerning L1 and L2 acquisition have looked almost exclusively at the first two factors. Those who believe that L1 learners have access to Universal Grammar whereas L2 learners rely on language processing strategies, postulate different learning mechanisms (UG parameter setting in L1, more general inductive strategies in L2 learning). Pienemann opposes this view and, based on his Processability Theory, argues that L1 and L2 learners start out from different initial states: they come to the grammar learning task with different structural hypotheses (SOV versus SVO as basic word order of German).
  • Kempen, G., Schotel, H., & Hoenkamp, E. (1982). Analyse-door-synthese van Nederlandse zinnen [Abstract]. De Psycholoog, 17, 509.
  • Kempen, G. (1995). 'Hier spreekt men Nederlands'. EMNET: Nieuwsbrief Elektronische Media, 22, 1.
  • Kempen, G. (1973). [Review of the book Psycholinguïstiek by B. Tervoort et al.]. Nederlands Tijdschrift voor de Psychologie, 28, 172-174.
  • Kempen, G., & Harbusch, K. (1998). A 'tree adjoining' grammar without adjoining: The case of scrambling in German. In Fourth International Workshop on Tree Adjoining Grammars and Related Frameworks (TAG+4).
  • Kempen, G. (1999). Fiets en (centri)fuge. Onze Taal, 68, 88.
  • Kempen, G. (1995). IJ of Y? Onze Taal, 64(9), 205-206.
  • Kempen, G., & Hoenkamp, E. (1982). Incremental sentence generation: Implications for the structure of a syntactic processor. In J. Horecký (Ed.), COLING 82. Proceedings of the Ninth International Conference on Computational Linguistics, Prague, July 5-10, 1982 (pp. 151-156). Amsterdam: North-Holland.

    Abstract

    Human speakers often produce sentences incrementally. They can start speaking having in mind only a fragmentary idea of what they want to say, and while saying this they refine the contents underlying subsequent parts of the utterance. This capability imposes a number of constraints on the design of a syntactic processor. This paper explores these constraints and evaluates some recent computational sentence generators from the perspective of incremental production.
  • Kempen, G., & De Vroomen, P. (Eds.). (1991). Informatiewetenschap 1991: Wetenschappelijke bijdragen aan de eerste STINFON-conferentie. Leiden: STINFON.
  • Kempen, G., & Sprangers, C. (Eds.). (1984). Kennis, mens en computer. Lisse: Swets & Zeitlinger.

    Abstract

    Essays van psychologen en linguı̈sten over de relatie hersens-computers.
  • Kempen, G. (1995). Processing discontinuous lexical items: A reply to Frazier. Cognition, 55, 219-221. doi:10.1016/0010-0277(94)00657-7.

    Abstract

    Comments on a study by Frazier and others on Dutch-language lexical processing. Claims that the control condition in the experiment was inadequate and that an assumption made by Frazier about closed class verbal items is inaccurate, and proposes an alternative account of a subset of the data from the experiment
  • Kempen, G. (1995). Processing separable complex verbs in Dutch: Comments on Frazier, Flores d'Arcais, and Coolen (1993). Cognition, 54, 353-356. doi:10.1016/0010-0277(94)00649-6.

    Abstract

    Raises objections to L. Frazier et al's (see record 1994-32229-001) report of an experimental study intended to test Schreuder's (1990) Morphological Integration (MI) model concerning the processing of separable and inseparable verbs and shows that the logic of the experiment is flawed. The problem is rooted in the notion of a separable complex verb. The conclusion is drawn that Frazier et al's experimental data cannot be taken as evidence for the theoretical propositions they develop about the MI model.
  • Kempen, G. (1998). Sentence parsing. In A. D. Friederici (Ed.), Language comprehension: A biological perspective (pp. 213-228). Berlin: Springer.
  • Kempen, G. (1984). Taaltechnologie voor het Nederlands: Vorderingen bij de bouw van een Nederlandstalig dialoog- en auteursysteem. Toegepaste Taalwetenschap in Artikelen, 19, 48-58.
  • Kempen, G., Konst, L., & De Smedt, K. (1984). Taaltechnologie voor het Nederlands: Vorderingen bij de bouw van een Nederlandstalig dialoog- en auteursysteem. Informatie, 26, 878-881.
  • Kempen, G. (1995). Van leescultuur en beeldcultuur naar internetcultuur. De Psycholoog, 30, 315-319.
  • Kempen, G. (1999). Visual Grammar: Multimedia for grammar and spelling instruction in primary education. In K. Cameron (Ed.), CALL: Media, design, and applications (pp. 223-238). Lisse: Swets & Zeitlinger.
  • Kendrick, K. H., Brown, P., Dingemanse, M., Floyd, S., Gipper, S., Hayano, K., Hoey, E., Hoymann, G., Manrique, E., Rossi, G., & Levinson, S. C. (2020). Sequence organization: A universal infrastructure for social action. Journal of Pragmatics, 168, 119-138. doi:10.1016/j.pragma.2020.06.009.

    Abstract

    This article makes the case for the universality of the sequence organization observable in informal human conversational interaction. Using the descriptive schema developed by Schegloff (2007), we examine the major patterns of action-sequencing in a dozen nearly all unrelated languages. What we find is that these patterns are instantiated in very similar ways for the most part right down to the types of different action sequences. There are also some notably different cultural exploitations of the patterns, but the patterns themselves look strongly universal. Recent work in gestural communication in the great apes suggests that sequence organization may have been a crucial route into the development of language. Taken together with the fundamental role of this organization in language acquisition, sequential behavior of this kind seems to have both phylogenetic and ontogenetic priority, which probably puts substantial functional pressure on language form.

    Additional information

    Supplementary data
  • Khan, M., & Cremers, F. P. (2020). ABCA4-Associated Stargardt Disease. Klinische Monatsblätter für Augenheilkunde, 237, 267-274. doi:10.1055/a-1057-9939.

    Abstract

    Autosomal recessive Stargardt disease (STGD1) is associated with variants in the ABCA4 gene. The phenotypes range from early-onset STGD1, that clinically resembles severe cone-rod dystrophy, to intermediate STGD1 and late-onset STGD1. These different phenotypes can be correlated with different combinations of ABCA4 variants which can be classified according to their degree of severity. A significant fraction of STGD1 cases, particularly late-onset STGD1 cases, were shown to carry only a single ABCA4 variant. A frequent coding variant (p.Asn1868Ile) was recently identified which – in combination with a severe ABCA4 variant – is generally associated with late-onset STGD1. In addition, an increasing number of rare deep-intronic variants have been found and some of these are also associated with late-onset STGD1. The effect of these and other variants on ABCA4 RNA was tested using in vitro assays in human kidney cells using specially designed midigenes. With stem cells and photoreceptor progenitor cells derived from patient skin or blood cells, retina-specific splice defects can be assessed. With expert clinical examination to distinguish STGD1 cases from other maculopathies, as well as in-depth genomics and transcriptomics data, it is now possible to identify both mutant ABCA4 alleles in > 95% of cases.
  • Khan, M., Cornelis, S. S., Sangermano, R., Post, I. J. M., Janssen Groesbeek, A., Amsu, J., Gilissen, C., Garanto, A., Collin, R. W. J., & Cremers, F. P. M. (2020). In or out? New insights on exon recognition through splice-site interdependency. International Journal of Molecular Sciences, 21: 2300. doi:10.3390/ijms21072300.

    Abstract

    Noncanonical splice-site mutations are an important cause of inherited diseases. Based on in vitro and stem-cell-based studies, some splice-site variants show a stronger splice defect than expected based on their predicted effects, suggesting that other sequence motifs influence the outcome. We investigated whether splice defects due to human-inherited-disease-associated variants in noncanonical splice-site sequences in ABCA4, DMD, and TMC1 could be rescued by strengthening the splice site on the other side of the exon. Noncanonical 5′- and 3′-splice-site variants were selected. Rescue variants were introduced based on an increase in predicted splice-site strength, and the effects of these variants were analyzed using in vitro splice assays in HEK293T cells. Exon skipping due to five variants in noncanonical splice sites of exons in ABCA4, DMD, and TMC1 could be partially or completely rescued by increasing the predicted strengths of the other splice site of the same exon. We named this mechanism “splicing interdependency”, and it is likely based on exon recognition by splicing machinery. Awareness of this interdependency is of importance in the classification of noncanonical splice-site variants associated with disease and may open new opportunities for treatments
  • Khoe, Y. H., Tsoukala, C., Kootstra, G. J., & Frank, S. L. (2020). Modeling cross-language structural priming in sentence production. In T. C. Stewart (Ed.), Proceedings of the 18th Annual Meeting of the International Conference on Cognitive Modeling (pp. 131-137). University Park, PA, USA: The Penn State Applied Cognitive Science Lab.

    Abstract

    A central question in the psycholinguistic study of multilingualism is how syntax is shared across languages. We implement a model to investigate whether error-based implicit learning can provide an account of cross-language structural priming. The model is based on the Dual-path model of
    sentence-production (Chang, 2002). We implement our model using the Bilingual version of Dual-path (Tsoukala, Frank, & Broersma, 2017). We answer two main questions: (1) Can structural priming of active and passive constructions occur between English and Spanish in a bilingual version of the Dual-
    path model? (2) Does cross-language priming differ quantitatively from within-language priming in this model? Our results show that cross-language priming does occur in the model. This finding adds to the viability of implicit learning as an account of structural priming in general and cross-language
    structural priming specifically. Furthermore, we find that the within-language priming effect is somewhat stronger than the cross-language effect. In the context of mixed results from
    behavioral studies, we interpret the latter finding as an indication that the difference between cross-language and within-
    language priming is small and difficult to detect statistically.
  • Kidd, E., & Donnelly, S. (2020). Individual differences in first language acquisition. Annual Review of Linguistics, 6, 319-340. doi:10.1146/annurev-linguistics-011619-030326.

    Abstract

    Humans vary in almost every dimension imaginable, and language is no
    exception. In this article, we review the past research that has focused on
    individual differences (IDs) in first language acquisition. We first consider
    how different theoretical traditions in language acquisition treat IDs, and
    we argue that a focus on IDs is important given its potential to reveal the
    developmental dynamics and architectural constraints of the linguistic system.
    We then review IDs research that has examined variation in children’s
    linguistic input, early speech perception, and vocabulary and grammatical
    development. In each case, we observe systematic and meaningful variation,
    such that variation in one domain (e.g., early auditory and speech
    processing) has meaningful developmental consequences for development
    in higher-order domains (e.g., vocabulary). The research suggests a high
    degree of integration across the linguistic system, in which development
    across multiple linguistic domains is tightly coupled.
  • Kidd, E., Bigood, A., Donnelly, S., Durrant, S., Peter, M. S., & Rowland, C. F. (2020). Individual differences in first language acquisition and their theoretical implications. In C. F. Rowland, A. L. Theakston, B. Ambridge, & K. E. Twomey (Eds.), Current Perspectives on Child Language Acquisition: How children use their environment to learn (pp. 189-219). Amsterdam: John Benjamins. doi:10.1075/tilar.27.09kid.

    Abstract

    Much of Lieven’s pioneering work has helped move the study of individual differences to the centre of child language research. The goal of the present chapter is to illustrate how the study of individual differences provides crucial insights into the language acquisition process. In part one, we summarise some of the evidence showing how pervasive individual differences are across the whole of the language system; from gestures to morphosyntax. In part two, we describe three causal factors implicated in explaining individual differences, which, we argue, must be built into any theory of language acquisition (intrinsic differences in the neurocognitive learning mechanisms, the child’s communicative environment, and developmental cascades in which each new linguistic skill that the child has to acquire depends critically on the prior acquisition of foundational abilities). In part three, we present an example study on the role of the speed of linguistic processing on vocabulary development, which illustrates our approach to individual differences. The results show evidence of a changing relationship between lexical processing speed and vocabulary over developmental time, perhaps as a result of the changing nature of the structure of the lexicon. The study thus highlights the benefits of an individual differences approach in building, testing, and constraining theories of language acquisition.
  • Kidd, E., Arciuli, J., Christiansen, M. H., Isbilen, E. S., Revius, K., & Smithson, M. (2020). Measuring children’s auditory statistical learning via serial recall. Journal of Experimental Child Psychology, 200: 104964. doi:10.1016/j.jecp.2020.104964.

    Abstract

    Statistical learning (SL) has been a prominent focus of research in developmental and adult populations, guided by the assumption that it is a fundamental component of learning underlying higher-order cognition. In developmental populations, however, there have been recent concerns regarding the degree to which many current tasks reliably measure SL, particularly in younger children. In the current article, we present the results of two studies that measured auditory statistical learning (ASL) of linguistic stimuli in children aged 5–8 years. Children listened to 6 min of continuous syllables comprising four trisyllabic pseudowords. Following the familiarization phase, children completed (a) a two-alternative forced-choice task and (b) a serial recall task in which they repeated either target sequences embedded during familiarization or foils, manipulated for sequence length. Results showed that, although both measures consistently revealed learning at the group level, the recall task better captured learning across the full range of abilities and was more reliable at the individual level. We conclude that, as has also been demonstrated in adults, the method holds promise for future studies of individual differences in ASL of linguistic stimuli.
  • Kim, N., Brehm, L., Sturt, P., & Yoshida, M. (2020). How long can you hold the filler: Maintenance and retrieval. Language, Cognition and Neuroscience, 35(1), 17-42. doi:10.1080/23273798.2019.1626456.

    Abstract

    This study attempts to reveal the mechanisms behind the online formation of Wh-Filler-Gap Dependencies (WhFGD). Specifically, we aim to uncover the way in which maintenance and retrieval work in WhFGD processing, by paying special attention to the information that is retrieved when the gap is recognized. We use the agreement attraction phenomenon (Wagers, M. W., Lau, E. F., & Phillips, C. (2009). Agreement attraction in comprehension: Representations and processes. Journal of Memory and Language, 61(2), 206-237) as a probe. The first and second experiments examined the type of information that is maintained and how maintenance is motivated, investigating the retrieved information at the gap for reactivated fillers and definite NPs. The third experiment examined the role of the retrieval, comparing reactivated and active fillers. We contend that the information being accessed reflects the extent to which the filler is maintained, where the reader is able to access fine-grained information including category information as well as a representation of both the head and the modifier at the verb.

    Additional information

    Supplemental material
  • Kita, S. (1995). Enter/exit animation for linguistic elicitation. In D. Wilkins (Ed.), Extensions of space and beyond: manual for field elicitation for the 1995 field season (pp. 13). Nijmegen: Max Planck Institute for Psycholinguistics. doi:10.17617/2.3003394.

    Abstract

    This task investigates the expression of “enter” and “exit” events, and is a supplement to the Motion Elicitation task (https://doi.org/10.17617/2.3003391). Consultants are asked to describe a series of animated clips where a man moves into or out of a house. The clips focus on contrasts to do with perspective (e.g., whether the man appears to move away or towards the viewer) and transitional movement (e.g., whether the man walks or “teleports” into his new location).

    Additional information

    1995_Enter_exit_animation_stimuli.zip
  • Kita, S., van Gijn, I., & van der Hulst, H. (1998). Movement phases in signs and co-speech gestures, and their transcription by human coders. In Gesture and Sign-Language in Human-Computer Interaction (Lecture Notes in Artificial Intelligence - LNCS Subseries, Vol. 1371) (pp. 23-35). Berlin, Germany: Springer-Verlag.

    Abstract

    The previous literature has suggested that the hand movement in co-speech gestures and signs consists of a series of phases with qualitatively different dynamic characteristics. In this paper, we propose a syntagmatic rule system for movement phases that applies to both co-speech gestures and signs. Descriptive criteria for the rule system were developed for the analysis video-recorded continuous production of signs and gesture. It involves segmenting a stream of body movement into phases and identifying different phase types. Two human coders used the criteria to analyze signs and cospeech gestures that are produced in natural discourse. It was found that the criteria yielded good inter-coder reliability. These criteria can be used for the technology of automatic recognition of signs and co-speech gestures in order to segment continuous production and identify the potentially meaningbearing phase.
  • Kita, S. (1995). Recommendations for data collection for gesture studies. In D. Wilkins (Ed.), Extensions of space and beyond: manual for field elicitation for the 1995 field season (pp. 35-45). Nijmegen: Max Planck Institute for Psycholinguistics. doi:10.17617/2.3004287.

    Abstract

    Do our hands 'speak the same language' across cultures? Gesture is the silent partner of spoken languages in face-to-face interaction, but we still have a lot to learn about gesture practices in different speech communities. The primary purpose of this task is to collect data in naturalistic settings that can be used to investigate the linguistic and cultural relativity of gesture performance, especially spatially indicative gestures. It involves video-recording pairs of speakers in both free conversation and more structured communication tasks (e.g., describing film plots).

    Please note: the stimuli mentioned in this entry are available elsewhere: 'The Pear Story', a short film made at the University of California at Berkeley; "Frog, where are you?" from the original Mayer (1969) book, as published in the Appendix of Berman & Slobin (1994).
  • Kita, S., & Ozyurek, A. (1999). Semantische Koordination zwischen Sprache und spontanen ikonischen Gesten: Eine sprachvergleichende Untersuchung. In Max-Planck-Gesellschaft (Ed.), Jahrbuch 1998 (pp. 388-391). Göttingen: Vandenhoeck & Ruprecht.
  • Klein, W. (1968). Zur Kategorisierung der Funktionsverbgefüge. Beiträge zur Linguistik und Informationsverarbeitung, 13, 7-37.
  • Klein, W. (1984). Bühler Ellipse. In C. F. Graumann, & T. Herrmann (Eds.), Karl Bühlers Axiomatik: Fünfzig Jahre Axiomatik der Sprachwissenschaften (pp. 117-141). Frankfurt am Main: Klostermann.
  • Klein, W. (1995). A simplest analysis of the English tense-aspect system. In W. Riehle, & H. Keiper (Eds.), Proceedings of the Anglistentag 1994 (pp. 139-151). Tübingen: Niemeyer.
  • Klein, W. (1995). A time-relational analysis of Russian aspect. Language, 71(4), 669-695.
  • Klein, W., Dietrich, R., & Noyau, C. (1995). Conclusions. In R. Dietrich, W. Klein, & C. Noyau (Eds.), The acquisition of temporality in a second language (pp. 261-280). Amsterdam: Benjamins.
  • Klein, W., & Musan, R. (Eds.). (1999). Das deutsche Perfekt [Special Issue]. Zeitschrift für Literaturwissenschaft und Linguistik, (113).
  • Klein, W. (1995). Das Vermächtnis der Geschichte, der Müll der Vergangenheit, oder: Wie wichtig ist zu wissen, was die Menschen früher getan oder geglaubt haben, für das, was wir jetzt tun oder glauben? Zeitschrift für Literaturwissenschaft und Linguistik; Metzler, Stuttgart, 100, 77-100.
  • Klein, W., & Rieck, B.-O. (1982). Der Erwerb der Personalpronomina im ungesteuerten Spracherwerb. Zeitschrift für Literaturwissenschaft und Linguistik, 45, 35-71.
  • Klein, W. (1998). Ein Blick zurück auf die Varietätengrammatik. In U. Ammon, K. Mattheier, & P. Nelde (Eds.), Sociolinguistica: Internationales Jahrbuch für europäische Soziolinguistik (pp. 22-38). Tübingen: Niemeyer.
  • Klein, W. (1973). Eine Analyse der Kerne in Schillers "Räuber". In S. Marcus (Ed.), Mathematische Poetik (pp. 326-333). Frankfurt am Main: Athenäum.
  • Klein, W. (1973). Eine Analyse der Kerne in Schillers "Räuber". Cahiers de linguistique théorique et appliquée, 10, 195-200.
  • Klein, W. (1982). Einige Bemerkungen zur Frageintonation. Deutsche Sprache, 4, 289-310.

    Abstract

    In the first, critical part of this study, a small sample of simple German sentences with their empirically determined pitch contours is used to demonstrate the incorrectness of numerous currently hold views of German sentence intonation. In the second, more constructive part, several interrogative sentence types are analysed and an attempt is made to show that intonation, besides other functions, indicates the permantently changing 'thematic score' in on-going discourse as well as certain validity claims.
  • Klein, W. (1982). Einleitung. Zeitschrift für Literaturwissenschaft und Linguistik; Metzler, Stuttgart, 12, 7-8.
  • Klein, W. (Ed.). (1995). Epoche [Special Issue]. Zeitschrift für Literaturwissenschaft und Linguistik, (100).
  • Klein, W. (1999). Die Lehren des Zweitspracherwerbs. In N. Dittmar, & A. Ramat (Eds.), Grammatik und Diskurs: Studien zum Erwerb des Deutschen und des Italienischen (pp. 279-290). Tübingen: Stauffenberg.
  • Klein, W. (1998). Assertion and finiteness. In N. Dittmar, & Z. Penner (Eds.), Issues in the theory of language acquisition: Essays in honor of Jürgen Weissenborn (pp. 225-245). Bern: Peter Lang.
  • Klein, W. (1973). Dialekt und Einheitssprache im Fremdsprachenunterricht. In Beiträge zu den Sommerkursen des Goethe-Instituts München (pp. 53-60).
  • Klein, W. (1995). Frame of analysis. In R. Dietrich, W. Klein, & C. Noyau (Eds.), The acquisition of temporality in a second language (pp. 17-29). Amsterdam: Benjamins.
  • Klein, W. (1991). Geile Binsenbüschel, sehr intime Gespielen: Ein paar Anmerkungen über Arno Schmidt als Übersetzer. Zeitschrift für Literaturwissenschaft und Linguistik, 84, 124-129.
  • Klein, W., & Weissenborn, J. (Eds.). (1982). Here and there: Cross-linguistic studies on deixis and demonstration. Amsterdam: Benjamins.
  • Klein, W. (1995). Literaturwissenschaft, Linguistik, LiLi. Zeitschrift für Literaturwissenschaft und Linguistik; Metzler, Stuttgart, (100), 1-10.
  • Klein, W. (1982). Local deixis in route directions. In R. Jarvella, & W. Klein (Eds.), Speech, place, and action: Studies in deixis and related topics (pp. 161-182). New York: Wiley.
  • Klein, W., & Von Stechow, A. (1982). Intonation und Bedeutung von Fokus. Konstanz: Universität Konstanz.
  • Klein, W. (Ed.). (1998). Kaleidoskop [Special Issue]. Zeitschrift für Literaturwissenschaft und Linguistik, (112).
  • Klein, W. (1982). Pronoms personnels et formes d'acquisition. Encrages, 8/9, 42-46.
  • Klein, W. (1991). Raumausdrücke. Linguistische Berichte, 132, 77-114.
  • Klein, W., & Von Stutterheim, C. (1991). Text structure and referential movement. Arbeitsberichte des Forschungsprogramms S&P: Sprache und Pragmatik, 22.
  • Klein, W. (Ed.). (1984). Textverständlichkeit - Textverstehen [Special Issue]. Zeitschrift für Literaturwissenschaft und Linguistik, (55).
  • Klein, W., Coenen, J., Van Helvert, K., & Hendriks, H. (1995). The acquisition of Dutch. In R. Dietrich, W. Klein, & C. Noyau (Eds.), The acquisition of temporality in a second language (pp. 117-143). Amsterdam: Benjamins.
  • Klein, W. (1995). The acquisition of English. In R. Dietrich, W. Klein, & C. Noyau (Eds.), The acquisition of temporality in a second language (pp. 31-70). Amsterdam: Benjamins.
  • Klein, W. (1998). The contribution of second language acquisition research. Language Learning, 48, 527-550. doi:10.1111/0023-8333.00057.

    Abstract

    During the last 25 years, second language acquisition (SLA) research hasmade considerable progress, but is still far from proving a solid basis for foreign language teaching, or from a general theory of SLA. In addition, its status within the linguistic disciplines is still very low. I argue this has not much to do with low empirical or theoretical standards in the field—in this regard, SLA research is fully competitive—but with a particular perspective on the acquisition process: SLA researches learners' utterances as deviations from a certain target, instead of genuine manifestations of underlying language capacity; it analyses them in terms of what they are not rather than what they are. For some purposes such a "target deviation perspective" makes sense, but it will not help SLA researchers to substantially and independently contribute to a deeper understanding of the structure and function of the human language faculty. Therefore, these findings will remain of limited interest to other scientists until SLA researchers consider learner varieties a normal, in fact typical, manifestation of this unique human capacity.
  • Klein, W. (1991). Seven trivia of language acquisition. In L. Eubank (Ed.), Point counterpoint: Universal grammar in the second language (pp. 49-70). Amsterdam: Benjamins.
  • Klein, W. (1991). SLA theory: Prolegomena to a theory of language acquisition and implications for Theoretical Linguistics. In T. Huebner, & C. Ferguson (Eds.), Crosscurrents in second language acquisition and linguistic theories (pp. 169-194). Amsterdam: Benjamins.
  • Klein, W. (Ed.). (1982). Speech, place, and action: Studies of language in context. New York: Wiley.
  • Klein, W. (1995). Sprachverhalten. In M. Amelang, & Pawlik (Eds.), Enzyklopädie der Psychologie (pp. 469-505). Göttingen: Hogrefe.
  • Klein, W., & Extra, G. (1982). Second language acquisition by adult immigrants: A European Science Foundation project. In R. E. V. Stuip, & W. Zwanenburg (Eds.), Handelingen van het zevenendertigste Nederlandse Filologencongres (pp. 127-136). Amsterdam: APA-Holland Universiteitspers.
  • Klein, W., & Vater, H. (1998). The perfect in English and German. In L. Kulikov, & H. Vater (Eds.), Typology of verbal categories: Papers presented to Vladimir Nedjalkov on the occasion of his 70th birthday (pp. 215-235). Tübingen: Niemeyer.
  • Klein, W. (1991). Was kann sich die Übersetzungswissenschaft von der Linguistik erwarten? Zeitschrift für Literaturwissenschaft und Linguistik, 84, 104-123.
  • Klein, W. (1999). Wie sich das deutsche Perfekt zusammensetzt. Zeitschrift für Literaturwissenschaft und Linguistik; Metzler, Stuttgart, (113), 52-85.
  • Klein, W. (Ed.). (1982). Zweitspracherwerb [Special Issue]. Zeitschrift für Literaturwissenschaft und Linguistik, (45).
  • Klein, W. (1984). Zweitspracherwerb: Eine Einführung. Königstein/TS: Athenäum.
  • Klein, W. (1998). Von der einfältigen Wißbegierde. Zeitschrift für Literaturwissenschaft und Linguistik, 112, 6-13.
  • Klenova, A. V., Goncharova, M. V., Kashentseva, T. A., & Naidenko, S. V. (2020). Voice breaking and its relation to body mass and testosterone level in the Siberian Crane (Leucogeranus leucogeranus). Journal of Ornithology, 161, 859-871. doi:10.1007/s10336-020-01773-w.

    Abstract

    Vocal development of cranes (Gruidae) has attracted scientifc interest due to its special stage, voice breaking. During voice breaking, chicks of diferent crane species produce calls with two fundamental frequencies that correspond to those in adult low-frequency and juvenile high-frequency vocalizations. However, triggers that afect voice breaking in cranes are mainly unknown. Here we studied the voice breaking in the Siberian Crane (Leucogeranus leucogeranus) and test its relation to the body mass and testosterone level. We analyzed 5846 calls, 39 body mass measurements and 60 blood samples from 11
    Siberian Crane chicks in 8 ages from 2.5 to 18 months of life together with 90 body mass measurements and 61 blood samples from 24 Siberian Crane adults. The individual duration of voice breaking and dates of its onset, culmination and completion depended neither on the body mass nor on the testosterone level at various ages. But we found correlation between the testosterone level and mean deltas of percentages of the high and low frequency components in Siberian Crane calls between the closest recording sessions. We also observed some coincidence in time between the mean dates of voice breaking onset and the termination of body mass gain (at 7.5 months of age), and between the mean dates of voice breaking completion and the start of a new breeding season. Similar relations have been shown previously for some other crane species. We also showed for the frst time that the mean dates of voice breaking culmination correlated with the signifcant increase of the testosterone level (at 10.5 months of age). So, we suggest that voice breaking in cranes may be triggered by the end of chicks’ body
    growth, is stimulated by the increase of testosterone level and ends soon after adult cranes stop taking care of their chicks.

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