Publications

Displaying 301 - 400 of 822
  • Kanero, J., Geçkin, V., Oranç, C., Mamus, E., Küntay, A. C., & Göksun, T. (2018). Social robots for early language learning: Current evidence and future directions. Child Development Perspectives, 12(3), 146-151. doi:10.1111/cdep.12277.

    Abstract

    In this article, we review research on child–robot interaction (CRI) to discuss how social robots can be used to scaffold language learning in young children. First we provide reasons why robots can be useful for teaching first and second languages to children. Then we review studies on CRI that used robots to help children learn vocabulary and produce language. The studies vary in first and second languages and demographics of the learners (typically developing children and children with hearing and communication impairments). We conclude that, although social robots are useful for teaching language to children, evidence suggests that robots are not as effective as human teachers. However, this conclusion is not definitive because robots that tutor students in language have not been evaluated rigorously and technology is advancing rapidly. We suggest that CRI offers an opportunity for research and list possible directions for that work.
  • Kanero, J., Franko, I., Oranç, C., Uluşahin, O., Koskulu, S., Adigüzel, Z., Küntay, A. C., & Göksun, T. (2018). Who can benefit from robots? Effects of individual differences in robot-assisted language learning. In Proceedings of the 8th International Conference on Development and Learning and Epigenetic Robotics (ICDL-EpiRob) (pp. 212-217). Piscataway, NJ, USA: IEEE.

    Abstract

    It has been suggested that some individuals may benefit more from social robots than do others. Using second
    language (L2) as an example, the present study examined how individual differences in attitudes toward robots and personality
    traits may be related to learning outcomes. Preliminary results with 24 Turkish-speaking adults suggest that negative attitudes
    toward robots, more specifically thoughts and anxiety about the negative social impact that robots may have on the society,
    predicted how well adults learned L2 words from a social robot. The possible implications of the findings as well as future directions are also discussed
  • Kapatsinski, V., & Harmon, Z. (2017). A Hebbian account of entrenchment and (over)-extension in language learning. In G. Gunzelmann, A. Howes, T. Tenbrink, & E. Davelaar (Eds.), Proceedings of the 39th Annual Meeting of the Cognitive Science Society (CogSci 2017) (pp. 2366-2371). Austin, TX: Cognitive Science Society.

    Abstract

    In production, frequently used words are preferentially extended to new, though related meanings. In comprehension, frequent exposure to a word instead makes the learner confident that all of the word’s legitimate uses have been experienced, resulting in an entrenched form-meaning mapping between the word and its experienced meaning(s). This results in a perception-production dissociation, where the forms speakers are most likely to map onto a novel meaning are precisely the forms that they believe can never be used that way. At first glance, this result challenges the idea of bidirectional form-meaning mappings, assumed by all current approaches to linguistic theory. In this paper, we show that bidirectional form-meaning mappings are not in fact challenged by this production-perception dissociation. We show that the production-perception dissociation is expected even if learners of the lexicon acquire simple symmetrical form-meaning associations through simple Hebbian learning.
  • Karadöller, D. Z., Sumer, B., & Ozyurek, A. (2017). Effects of delayed language exposure on spatial language acquisition by signing children and adults. In G. Gunzelmann, A. Howes, T. Tenbrink, & E. Davelaar (Eds.), Proceedings of the 39th Annual Conference of the Cognitive Science Society (CogSci 2017) (pp. 2372-2376). Austin, TX: Cognitive Science Society.

    Abstract

    Deaf children born to hearing parents are exposed to language input quite late, which has long-lasting effects on language production. Previous studies with deaf individuals mostly focused on linguistic expressions of motion events, which have several event components. We do not know if similar effects emerge in simple events such as descriptions of spatial configurations of objects. Moreover, previous data mainly come from late adult signers. There is not much known about language development of late signing children soon after learning sign language. We compared simple event descriptions of late signers of Turkish Sign Language (adults, children) to age-matched native signers. Our results indicate that while late signers in both age groups are native-like in frequency of expressing a relational encoding, they lag behind native signers in using morphologically complex linguistic forms compared to other simple forms. Late signing children perform similar to adults and thus showed no development over time.
  • Kavaklioglu, T., Guadalupe, T., Zwiers, M., Marquand, A. F., Onnink, M., Shumskaya, E., Brunner, H., Fernandez, G., Fisher, S. E., & Francks, C. (2017). Structural asymmetries of the human cerebellum in relation to cerebral cortical asymmetries and handedness. Brain Structure and Function, 22, 1611-1623. doi:10.1007/s00429-016-1295-9.

    Abstract

    There is evidence that the human cerebellum is involved not only in motor control but also in other cognitive functions. Several studies have shown that language-related activation is lateralized toward the right cerebellar hemisphere in most people, in accordance with leftward cerebral cortical lateralization for language and a general contralaterality of cerebral–cerebellar activations. In terms of behavior, hand use elicits asymmetrical activation in the cerebellum, while hand preference is weakly associated with language lateralization. However, it is not known how, or whether, these functional relations are reflected in anatomy. We investigated volumetric gray matter asymmetries of cerebellar lobules in an MRI data set comprising 2226 subjects. We tested these cerebellar asymmetries for associations with handedness, and for correlations with cerebral cortical anatomical asymmetries of regions important for language or hand motor control, as defined by two different automated image analysis methods and brain atlases, and supplemented with extensive visual quality control. No significant associations of cerebellar asymmetries to handedness were found. Some significant associations of cerebellar lobular asymmetries to cerebral cortical asymmetries were found, but none of these correlations were greater than 0.14, and they were mostly method-/atlas-dependent. On the basis of this large and highly powered study, we conclude that there is no overt structural manifestation of cerebellar functional lateralization and connectivity, in respect of hand motor control or language laterality
  • Kember, H., Grohe, A.-.-K., Zahner, K., Braun, B., Weber, A., & Cutler, A. (2017). Similar prosodic structure perceived differently in German and English. In Proceedings of Interspeech 2017 (pp. 1388-1392). doi:10.21437/Interspeech.2017-544.

    Abstract

    English and German have similar prosody, but their speakers realize some pitch falls (not rises) in subtly different ways. We here test for asymmetry in perception. An ABX discrimination task requiring F0 slope or duration judgements on isolated vowels revealed no cross-language difference in duration or F0 fall discrimination, but discrimination of rises (realized similarly in each language) was less accurate for English than for German listeners. This unexpected finding may reflect greater sensitivity to rising patterns by German listeners, or reduced sensitivity by English listeners as a result of extensive exposure to phrase-final rises (“uptalk”) in their language
  • Kempen, G., & Harbusch, K. (2018). A competitive mechanism selecting verb-second versus verb-final word order in causative and argumentative clauses of spoken Dutch: A corpus-linguistic study. Language Sciences, 69, 30-42. doi:10.1016/j.langsci.2018.05.005.

    Abstract

    In Dutch and German, the canonical order of subject, object(s) and finite verb is ‘verb-second’ (V2) in main but ‘verb-final’ (VF) in subordinate clauses. This occasionally leads to the production of noncanonical word orders. Familiar examples are causative and argumentative clauses introduced by a subordinating conjunction (Du. omdat, Ger. weil ‘because’): the omdat/weil-V2 phenomenon. Such clauses may also be introduced by coordinating conjunctions (Du. want, Ger. denn), which license V2 exclusively. However, want/denn-VF structures are unknown. We present the results of a corpus study on the incidence of omdat-V2 in spoken Dutch, and compare them to published data on weil-V2 in spoken German. Basic findings: omdat-V2 is much less frequent than weil-V2 (ratio almost 1:8); and the frequency relations between coordinating and subordinating conjunctions are opposite (want >> omdat; denn << weil). We propose that conjunction selection and V2/VF selection proceed partly independently, and sometimes miscommunicate—e.g. yielding omdat/weil paired with V2. Want/denn-VF pairs do not occur because want/denn clauses are planned as autonomous sentences, which take V2 by default. We sketch a simple feedforward neural network with two layers of nodes (representing conjunctions and word orders, respectively) that can simulate the observed data pattern through inhibition-based competition of the alternative choices within the node layers.
  • Kempen, G. (1976). De taalgebruiker in de mens: Een uitzicht over de taalpsychologie. Groningen: H.D. Tjeenk Willink.
  • Kempen, G., Schotel, H., & Hoenkamp, E. (1982). Analyse-door-synthese van Nederlandse zinnen [Abstract]. De Psycholoog, 17, 509.
  • Kempen, G. (1973). [Review of the book Psycholinguïstiek by B. Tervoort et al.]. Nederlands Tijdschrift voor de Psychologie, 28, 172-174.
  • Kempen, G., & Harbusch, K. (2017). Frequential test of (S)OV as unmarked word order in Dutch and German clauses: A serendipitous corpus-linguistic experiment. In H. Reckman, L. L. S. Cheng, M. Hijzelendoorn, & R. Sybesma (Eds.), Crossroads semantics: Computation, experiment and grammar (pp. 107-123). Amsterdam: Benjamins.

    Abstract

    In a paper entitled “Against markedness (and what to replace it with)”, Haspelmath argues “that the term ‘markedness’ is superfluous”, and that frequency asymmetries often explain structural (un)markedness asymmetries (Haspelmath 2006). We investigate whether this argument applies to Object and Verb orders in main (VO, marked) and subordinate (OV, unmarked) clauses of spoken and written German and Dutch, using English (without VO/OV alternation) as control. Frequency counts from six treebanks (three languages, two output modalities) do not support Haspelmath’s proposal. However, they reveal an unexpected phenomenon, most prominently in spoken Dutch and German: a small set of extremely high-frequent finite verbs with unspecific meanings populates main clauses much more densely than subordinate clauses. We suggest these verbs accelerate the start-up of grammatical encoding, thus facilitating sentence-initial output fluency
  • Kempen, G., & Hoenkamp, E. (1982). Incremental sentence generation: Implications for the structure of a syntactic processor. In J. Horecký (Ed.), COLING 82. Proceedings of the Ninth International Conference on Computational Linguistics, Prague, July 5-10, 1982 (pp. 151-156). Amsterdam: North-Holland.

    Abstract

    Human speakers often produce sentences incrementally. They can start speaking having in mind only a fragmentary idea of what they want to say, and while saying this they refine the contents underlying subsequent parts of the utterance. This capability imposes a number of constraints on the design of a syntactic processor. This paper explores these constraints and evaluates some recent computational sentence generators from the perspective of incremental production.
  • Kempen, G. (1976). Syntactic constructions as retrieval plans. British Journal of Psychology, 67(2), 149-160. doi:10.1111/j.2044-8295.1976.tb01505.x.

    Abstract

    Four probe latency experiments show that the ‘constituent boundary effect’ (transitions between constituents are more difficult than within constituents) is a retrieval and not a storage phenomenon. The experimental logic used is called paraphrastic reproduction: after verbatim memorization of some sentences, subjects were instructed to reproduce them both in their original wording and in the form of sentences that, whilst preserving the original meaning, embodied different syntactic constructions. Syntactic constructions are defined as pairs which consist of a pattern of conceptual information and a syntactic scheme, i.e. a sequence of syntactic word categories and function words. For example, the sequence noun + finite intransitive main verb (‘John runs’) expresses a conceptual actor-action relationship. It is proposed that for each overlearned and simple syntactic construction there exists a retrieval plan which does the following. It searches through the long-term memory information that has been designated as the conceptual content of the utterance(s) to be produced, looking for a token of its conceptual pattern. The retrieved information is then cast into the format of its syntactic scheme. The organization of such plans is held responsible for the constituent boundary effect.
  • Kendrick, K. H., & Holler, J. (2017). Gaze direction signals response preference in conversation. Research on Language and Social Interaction, 50(1), 12-32. doi:10.1080/08351813.2017.1262120.

    Abstract

    In this article, we examine gaze direction in responses to polar questions using both quantitative and conversation analytic (CA) methods. The data come from a novel corpus of conversations in which participants wore eye-tracking glasses to obtain direct measures of their eye movements. The results show that while most preferred responses are produced with gaze toward the questioner, most dispreferred responses are produced with gaze aversion. We further demonstrate that gaze aversion by respondents can occasion self-repair by questioners in the transition space between turns, indicating that the relationship between gaze direction and preference is more than a mere statistical association. We conclude that gaze direction in responses to polar questions functions as a signal of response preference. Data are in American, British, and Canadian English.

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  • Ketrez, F. N., Kuntay, A. C., Ozcaliskan, S., & Ozyurek, A. (Eds.). (2017). Social environment and cognition in language development: Studies in honor of Ayhan Aksu-Koc. Amsterdam: John Benjamins.

    Abstract

    Language development is driven by multiple factors involving both the individual child and the environments that surround the child. The chapters in this volume highlight several such factors as potential contributors to developmental change, including factors that examine the role of immediate social environment (i.e., parent SES, parent and sibling input, peer interaction) and factors that focus on the child’s own cognitive and social development, such as the acquisition of theory of mind, event knowledge, and memory. The discussion of the different factors is presented largely from a crosslinguistic framework, using a multimodal perspective (speech, gesture, sign). The book celebrates the scholarly contributions of Prof. Ayhan Aksu-Koç – a pioneer in the study of crosslinguistic variation in language acquisition, particularly in the domain of evidentiality and theory of mind. This book will serve as an important resource for researchers in the field of developmental psychology, cognitive science, and linguistics across the globe
  • Kidd, E., Junge, C., Spokes, T., Morrison, L., & Cutler, A. (2018). Individual differences in infant speech segmentation: Achieving the lexical shift. Infancy, 23(6), 770-794. doi:10.1111/infa.12256.

    Abstract

    We report a large‐scale electrophysiological study of infant speech segmentation, in which over 100 English‐acquiring 9‐month‐olds were exposed to unfamiliar bisyllabic words embedded in sentences (e.g., He saw a wild eagle up there), after which their brain responses to either the just‐familiarized word (eagle) or a control word (coral) were recorded. When initial exposure occurs in continuous speech, as here, past studies have reported that even somewhat older infants do not reliably recognize target words, but that successful segmentation varies across children. Here, we both confirm and further uncover the nature of this variation. The segmentation response systematically varied across individuals and was related to their vocabulary development. About one‐third of the group showed a left‐frontally located relative negativity in response to familiar versus control targets, which has previously been described as a mature response. Another third showed a similarly located positive‐going reaction (a previously described immature response), and the remaining third formed an intermediate grouping that was primarily characterized by an initial response delay. A fine‐grained group‐level analysis suggested that a developmental shift to a lexical mode of processing occurs toward the end of the first year, with variation across individual infants in the exact timing of this shift.

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  • Kidd, E., Donnelly, S., & Christiansen, M. H. (2018). Individual differences in language acquisition and processing. Trends in Cognitive Sciences, 22(2), 154-169. doi:10.1016/j.tics.2017.11.006.

    Abstract

    Humans differ in innumerable ways, with considerable variation observable at every level of description, from the molecular to the social. Traditionally, linguistic and psycholinguistic theory has downplayed the possibility of meaningful differences in language across individuals. However, it is becoming increasingly evident that there is
    significant variation among speakers at any age as well as across the lifespan. In this paper, we review recent research in psycholinguistics, and argue that a focus on individual differences provides a crucial source of evidence that bears strongly upon core issues in theories of the acquisition and processing of language; specifically, the role of experience in language acquisition, processing, and attainment, and the architecture of the language faculty.
  • Kirjavainen, M., Kidd, E., & Lieven, E. (2017). How do language-specific characteristics affect the acquisition of different relative clause types? Evidence from Finnish. Journal of Child Language, 44(1), 120-157. doi:10.1017/s0305000915000768.

    Abstract

    We report three studies (one corpus, two experimental) that investigated the acquisition of relative clauses (RCs) in Finnish-speaking children. Study 1 found that Finnish children's naturalistic exposure to RCs predominantly consists of non-subject relatives (i.e. oblique, object) which typically have inanimate head nouns. Study 2 tested children's comprehension of subject, object, and two types of oblique relatives. No difference was found in the children's performance on different structures, including a lack of previously widely reported asymmetry between subject and object relatives. However, children's comprehension was modulated by animacy of the head referent. Study 3 tested children's production of the same RC structures using sentence repetition. Again we found no subject–object asymmetry. The pattern of results suggested that distributional frequency patterns and the relative complexity of the relativizer contribute to the difficulty associated with particular RC structures.
  • Kirkham, J. A., & Kidd, E. (2017). The effect of Steiner, Montessori, and national curriculum education upon children's pretence and creativity. Journal of Creative Behavior, 51(1), 20-34. doi:10.1002/jocb.83.

    Abstract

    Pretence and creativity are often regarded as ubiquitous characteristics of childhood, yet not all education systems value or promote these attributes to the same extent. Different pedagogies and practices are evident within the UK National Curriculum, Steiner and Montessori schools. In this study, 20 children participated from each of these school systems (N = 60, aged 6;10–8;11) completing the test of creative thinking—drawing production (TCT-DP; K. K. Urban, & H. G. Jellen, 1996) and a pretend actions task (W. F. Overton & J. P. Jackson, 1973). Overall, Steiner pupils performed significantly higher on the TCT-DP than both the Montessori and National Curriculum pupils who performed similarly. Steiner pupils also performed significantly better on the pretend actions task than the Montessori pupils, but no other significant differences were found. Overall, there was also a significant positive correlation between pretence and creativity in the current sample, supporting previous research suggesting that these skills are related (e.g., A.S. Kaugars & S. W. Russ, 2009; P. Y. Mullineaux & L. F. Dilalla, 2009).
  • Kirsch, J. (2018). Listening for the WHAT and the HOW: Older adults' processing of semantic and affective information in speech. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Kita, S., Alibali, M. W., & Chu, M. (2017). How Do Gestures Influence Thinking and Speaking? The Gesture-for-Conceptualization Hypothesis. Psychological Review, 124(3), 245-266. doi:10.1037/rev0000059.

    Abstract

    People spontaneously produce gestures during speaking and thinking. The authors focus here on gestures that depict or indicate information related to the contents of concurrent speech or thought (i.e., representational gestures). Previous research indicates that such gestures have not only communicative functions, but also self-oriented cognitive functions. In this article, the authors propose a new theoretical framework, the gesture-for-conceptualization hypothesis, which explains the self-oriented functions of representational gestures. According to this framework, representational gestures affect cognitive processes in 4 main ways: gestures activate, manipulate, package, and explore spatio-motoric information for speaking and thinking. These four functions are shaped by gesture's ability to schematize information, that is, to focus on a small subset of available information that is potentially relevant to the task at hand. The framework is based on the assumption that gestures are generated from the same system that generates practical actions, such as object manipulation; however, gestures are distinct from practical actions in that they represent information. The framework provides a novel, parsimonious, and comprehensive account of the self-oriented functions of gestures. The authors discuss how the framework accounts for gestures that depict abstract or metaphoric content, and they consider implications for the relations between self-oriented and communicative functions of gestures
  • Kiyama, S., Verdonschot, R. G., Xiong, K., & Tamaoka, K. (2018). Individual mentalizing ability boosts flexibility toward a linguistic marker of social distance: An ERP investigation. Journal of Neurolinguistics, 47, 1-15. doi:10.1016/j.jneuroling.2018.01.005.

    Abstract

    Sentence-final particles (SFPs) as bound morphemes in Japanese have no obvious effect on the truth conditions of a sentence. However, they encompass a diverse range of usages, from typical to atypical, according to the context and the interpersonal relationships in the specific situation. The most frequent particle,-ne, is typically used after addressee-oriented propositions for information sharing, while another frequent particle,-yo, is typically used after addresser-oriented propositions to elicit a sense of strength. This study sheds light on individual differences among native speakers in flexibly understanding such linguistic markers based on their mentalizing ability (i.e., the ability to infer the mental states of others). Two experiments employing electroencephalography (EEG) consistently showed enhanced early posterior negativities (EPN) for atypical SFP usage compared to typical usage, especially when understanding-ne compared to -yo, in both an SFP appropriateness judgment task and a content comprehension task. Importantly, the amplitude of the EPN for atypical usages of-ne was significantly higher in participants with lower mentalizing ability than in those with a higher mentalizing ability. This effect plausibly reflects low-ability mentalizers' stronger sense of strangeness toward atypical-ne usage. While high-ability mentalizers may aptly perceive others' attitudes via their various usages of-ne, low-ability mentalizers seem to adopt a more stereotypical understanding. These results attest to the greater degree of difficulty low-ability mentalizers have in establishing a smooth regulation of interpersonal distance during social encounters.

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  • Klamer, M., Trilsbeek, P., Hoogervorst, T., & Haskett, C. (2017). Creating a Language Archive of Insular South East Asia and West New Guinea. In J. Odijk, & A. Van Hessen (Eds.), CLARIN in the Low Countries (pp. 113-121). London: Ubiquity Press. doi:10.5334/bbi.10.

    Abstract

    The geographical region of Insular South East Asia and New Guinea is well-known as an
    area of mega-biodiversity. Less well-known is the extreme linguistic diversity in this area:
    over a quarter of the world’s 6,000 languages are spoken here. As small minority languages,
    most of them will cease to be spoken in the coming few generations. The project described
    here ensures the preservation of unique records of languages and the cultures encapsulated
    by them in the region. The language resources were gathered by twenty linguists at,
    or in collaboration with, Dutch universities over the last 40 years, and were compiled and
    archived in collaboration with The Language Archive (TLA) at the Max Planck Institute in
    Nijmegen. The resulting archive constitutes a collection ofmultimediamaterials and written
    documents from 48 languages in Insular South East Asia and West New Guinea. At TLA,
    the data was archived according to state-of-the-art standards (TLA holds the Data Seal of
    Approval): the component metadata infrastructure CMDI was used; all metadata categories
    as well as relevant units of annotation were linked to the ISO data category registry ISOcat.
    This guaranteed proper integration of the language resources into the CLARIN framework.
    Through the archive, future speaker communities and researchers will be able to extensively
    search thematerials for answers to their own questions, even if they do not themselves know the language, and even if the language dies.
  • Klein, W. (2017). Statt einer Einleitung: Brigitte Schlieben-Lange zum Gedenken. Zeitschrift für Literaturwissenschaft und Linguistik, 30(4), 5-10. doi:10.1007/BF03379240.
  • Klein, W., & Rieck, B.-O. (1982). Der Erwerb der Personalpronomina im ungesteuerten Spracherwerb. Zeitschrift für Literaturwissenschaft und Linguistik, 45, 35-71.
  • Klein, W. (1976). Der Prozeß des Zweitspracherwerbs und seine Beschreibung. In R. Dietrich (Ed.), Aspekte des Fremdsprachenerwerbs (pp. 100-118). Kronberg/Ts.: Athenäum.
  • Klein, W. (1973). Eine Analyse der Kerne in Schillers "Räuber". In S. Marcus (Ed.), Mathematische Poetik (pp. 326-333). Frankfurt am Main: Athenäum.
  • Klein, W. (1973). Eine Analyse der Kerne in Schillers "Räuber". Cahiers de linguistique théorique et appliquée, 10, 195-200.
  • Klein, W. (1982). Einige Bemerkungen zur Frageintonation. Deutsche Sprache, 4, 289-310.

    Abstract

    In the first, critical part of this study, a small sample of simple German sentences with their empirically determined pitch contours is used to demonstrate the incorrectness of numerous currently hold views of German sentence intonation. In the second, more constructive part, several interrogative sentence types are analysed and an attempt is made to show that intonation, besides other functions, indicates the permantently changing 'thematic score' in on-going discourse as well as certain validity claims.
  • Klein, W. (1976). Einige wesentliche Eigenschaften natürlicher Sprachen und ihre Bedeutung für die linguistische Theorie. Zeitschrift für Literaturwissenschaft und Linguistik, 23/24, 11-31.
  • Klein, W. (1976). Einleitung. Zeitschrift für Literaturwissenschaft und Linguistik; Metzler, Stuttgart, 6(23/24), 7-10.
  • Klein, W. (1982). Einleitung. Zeitschrift für Literaturwissenschaft und Linguistik; Metzler, Stuttgart, 12, 7-8.
  • Klein, W. (1973). Dialekt und Einheitssprache im Fremdsprachenunterricht. In Beiträge zu den Sommerkursen des Goethe-Instituts München (pp. 53-60).
  • Klein, M., Van Donkelaar, M., Verhoef, E., & Franke, B. (2017). Imaging genetics in neurodevelopmental psychopathology. American Journal of Medical Genetics Part B: Neuropsychiatric Genetics, 174(5), 485-537. doi:10.1002/ajmg.b.32542.

    Abstract

    Neurodevelopmental disorders are defined by highly heritable problems during development and brain growth. Attention-deficit/hyperactivity disorder (ADHD), autism spectrum disorders (ASDs), and intellectual disability (ID) are frequent neurodevelopmental disorders, with common comorbidity among them. Imaging genetics studies on the role of disease-linked genetic variants on brain structure and function have been performed to unravel the etiology of these disorders. Here, we reviewed imaging genetics literature on these disorders attempting to understand the mechanisms of individual disorders and their clinical overlap. For ADHD and ASD, we selected replicated candidate genes implicated through common genetic variants. For ID, which is mainly caused by rare variants, we included genes for relatively frequent forms of ID occurring comorbid with ADHD or ASD. We reviewed case-control studies and studies of risk variants in healthy individuals. Imaging genetics studies for ADHD were retrieved for SLC6A3/DAT1, DRD2, DRD4, NOS1, and SLC6A4/5HTT. For ASD, studies on CNTNAP2, MET, OXTR, and SLC6A4/5HTT were found. For ID, we reviewed the genes FMR1, TSC1 and TSC2, NF1, and MECP2. Alterations in brain volume, activity, and connectivity were observed. Several findings were consistent across studies, implicating, for example, SLC6A4/5HTT in brain activation and functional connectivity related to emotion regulation. However, many studies had small sample sizes, and hypothesis-based, brain region-specific studies were common. Results from available studies confirm that imaging genetics can provide insight into the link between genes, disease-related behavior, and the brain. However, the field is still in its early stages, and conclusions about shared mechanisms cannot yet be drawn.
  • Klein, W. (1976). Maschinelle Analyse des Sprachwandels. In P. Eisenberg (Ed.), Maschinelle Sprachanalyse (pp. 137-166). Berlin: de Gruyter.
  • Klein, W., & Weissenborn, J. (Eds.). (1982). Here and there: Cross-linguistic studies on deixis and demonstration. Amsterdam: Benjamins.
  • Klein, W. (1982). Local deixis in route directions. In R. Jarvella, & W. Klein (Eds.), Speech, place, and action: Studies in deixis and related topics (pp. 161-182). New York: Wiley.
  • Klein, W. (2018). Looking at language. Berlin: De Gruyter.

    Abstract

    The volume presents an essential selection collected from the essays of Wolfgang Klein. In addition to journal and book articles, many of them published by Mouton, this book features new and unpublished texts by the author. It focuses, among other topics, on information structure, the expression of grammatical categories and the structure of learner varieties.
  • Klein, W., & Von Stechow, A. (1982). Intonation und Bedeutung von Fokus. Konstanz: Universität Konstanz.
  • Klein, W. (1982). Pronoms personnels et formes d'acquisition. Encrages, 8/9, 42-46.
  • Klein, W. (Ed.). (1976). Psycholinguistik [Special Issue]. Zeitschrift für Literaturwissenschaft und Linguistik, (23/24).
  • Klein, W. (Ed.). (1982). Speech, place, and action: Studies of language in context. New York: Wiley.
  • Klein, W. (1976). Sprachliche Variation. Studium Linguistik, 1, 29-46.
  • Klein, W., & Extra, G. (1982). Second language acquisition by adult immigrants: A European Science Foundation project. In R. E. V. Stuip, & W. Zwanenburg (Eds.), Handelingen van het zevenendertigste Nederlandse Filologencongres (pp. 127-136). Amsterdam: APA-Holland Universiteitspers.
  • Klein, W. (Ed.). (1982). Zweitspracherwerb [Special Issue]. Zeitschrift für Literaturwissenschaft und Linguistik, (45).
  • Klepp, A., Niccolai, V., Sieksmeyer, J., Arnzen, S., Indefrey, P., Schnitzler, A., & Biermann-Ruben, K. (2017). Body-part specific interactions of action verb processing with motor behaviour. Behavioural Brain Research, 328, 149-158. doi:10.1016/j.bbr.2017.04.002.

    Abstract

    The interaction of action-related language processing with actual movement is an indicator of the functional role of motor cortical involvement in language understanding. This paper describes two experiments using single action verb stimuli. Motor responses were performed with the hand or the foot. To test the double dissociation of language-motor facilitation effects within subjects, Experiments 1 and 2 used a priming procedure where both hand and foot reactions had to be performed in response to different geometrical shapes, which were preceded by action verbs. In Experiment 1, the semantics of the verbs could be ignored whereas Experiment 2 included semantic decisions. Only Experiment 2 revealed a clear double dissociation in reaction times: reactions were facilitated when preceded by verbs describing actions with the matching effector. In Experiment 1, by contrast, there was an interaction between verb-response congruence and a semantic variable related to motor features of the verbs. Thus, the double dissociation paradigm of semantic motor priming was effective, corroborating the role of the motor system in action-related language processing. Importantly, this effect was body part specific.

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  • Koch, X. (2018). Age and hearing loss effects on speech processing. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Kochari, A. R., & Ostarek, M. (2018). Introducing a replication-first rule for PhD projects (commmentary on Zwaan et al., ‘Making replication mainstream’). Behavioral and Brain Sciences, 41: e138. doi:10.1017/S0140525X18000730.

    Abstract

    Zwaan et al. mention that young researchers should conduct replications as a
    small part of their portfolio. We extend this proposal and suggest that conducting and
    reporting replications should become an integral part of PhD projects and be taken into
    account in their assessment. We discuss how this would help not only scientific
    advancement, but also PhD candidates’ careers.
  • Kolipakam, V. (2018). A holistic approach to understanding pre-history. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Kolipakam, V., Jordan, F., Dunn, M., Greenhill, S. J., Bouckaert, R., Gray, R. D., & Verkerk, A. (2018). A Bayesian phylogenetic study of the Dravidian language family. Royal Society Open Science, 5: 171504. doi:10.1098/rsos.171504.

    Abstract

    The Dravidian language family consists of about 80 varieties (Hammarström H. 2016 Glottolog 2.7) spoken by 220 million people across southern and central India and surrounding countries (Steever SB. 1998 In The Dravidian languages (ed. SB Steever), pp. 1–39: 1). Neither the geographical origin of the Dravidian language homeland nor its exact dispersal through time are known. The history of these languages is crucial for understanding prehistory in Eurasia, because despite their current restricted range, these languages played a significant role in influencing other language groups including Indo-Aryan (Indo-European) and Munda (Austroasiatic) speakers. Here, we report the results of a Bayesian phylogenetic analysis of cognate-coded lexical data, elicited first hand from native speakers, to investigate the subgrouping of the Dravidian language family, and provide dates for the major points of diversification. Our results indicate that the Dravidian language family is approximately 4500 years old, a finding that corresponds well with earlier linguistic and archaeological studies. The main branches of the Dravidian language family (North, Central, South I, South II) are recovered, although the placement of languages within these main branches diverges from previous classifications. We find considerable uncertainty with regard to the relationships between the main branches.
  • Kong, X., Song, Y., Zhen, Z., & Liu, J. (2017). Genetic Variation in S100B Modulates Neural Processing of Visual Scenes in Han Chinese. Cerebral Cortex, 27(2), 1326-1336. doi:10.1093/cercor/bhv322.

    Abstract

    Spatial navigation is a crucial ability for living. Previous animal studies have shown that the S100B gene is causally related to spatial navigation performance in mice. However, the genetic factors influencing human navigation and its neural substrates remain unclear. Here, we provided the first evidence that the S100B gene modulates neural processing of navigationally relevant scenes in humans. First, with a novel protocol, we demonstrated that the spatial pattern of S100B gene expression in postmortem brains was associated with brain activation pattern for spatial navigation in general, and for scene processing in particular. Further, in a large fMRI cohort of healthy adults of Han Chinese (N = 202), we found that S100B gene polymorphisms modulated scene selectivity in the retrosplenial cortex (RSC) and parahippocampal place area. Finally, the serum levels of S100B protein mediated the association between S100B gene polymorphism and scene selectivity in the RSC. Our study takes the first step toward understanding the neurogenetic mechanism of human spatial navigation and suggests a novel approach to discover candidate genes modulating cognitive functions.

    Additional information

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  • Kong, X., Wang, X., Pu, Y., Huang, L., Hao, X., Zhen, Z., & Liu, J. (2017). Human navigation network: The intrinsic functional organization and behavioral relevance. Brain Structure and Function, 222(2), 749-764. doi:10.1007/s00429-016-1243-8.

    Abstract

    Spatial navigation is a crucial ability for living. Previous work has revealed multiple distributed brain regions associated with human navigation. However, little is known about how these regions work together as a network (referred to as navigation network) to support flexible navigation. In a novel protocol, we combined neuroimaging meta-analysis, and functional connectivity and behavioral data from the same subjects. Briefly, we first constructed the navigation network for each participant, by combining a large-scale neuroimaging meta-analysis (with the Neurosynth) and resting-state functional magnetic resonance imaging. Then, we investigated multiple topological properties of the navigation networks, including small-worldness, modularity, and highly connected hubs. Finally, we explored the behavioral relevance of these intrinsic properties in a large sample of healthy young adults (N = 190). We found that navigation networks showed small-world and modular organization at global level. More importantly, we found that increased small-worldness and modularity of the navigation network were associated with better navigation ability. Finally, we found that the right retrosplenial complex (RSC) acted as one of the hubs in the navigation network, and that higher betweenness of this region correlated with better navigation ability, suggesting a critical role of the RSC in modulating the navigation network in human brain. Our study takes one of the first steps toward understanding the underlying organization of the navigation network. Moreover, these findings suggest the potential applications of the novel approach to investigating functionally meaningful networks in human brain and their relations to the behavioral impairments in the aging and psychiatric patients.
  • Kong, X., Mathias, S. R., Guadalupe, T., ENIGMA Laterality Working Group, Glahn, D. C., Franke, B., Crivello, F., Tzourio-Mazoyer, N., Fisher, S. E., Thompson, P. M., & Francks, C. (2018). Mapping Cortical Brain Asymmetry in 17,141 Healthy Individuals Worldwide via the ENIGMA Consortium. Proceedings of the National Academy of Sciences of the United States of America, 115(22), E5154-E5163. doi:10.1073/pnas.1718418115.

    Abstract

    Hemispheric asymmetry is a cardinal feature of human brain organization. Altered brain asymmetry has also been linked to some cognitive and neuropsychiatric disorders. Here the ENIGMA consortium presents the largest ever analysis of cerebral cortical asymmetry and its variability across individuals. Cortical thickness and surface area were assessed in MRI scans of 17,141 healthy individuals from 99 datasets worldwide. Results revealed widespread asymmetries at both hemispheric and regional levels, with a generally thicker cortex but smaller surface area in the left hemisphere relative to the right. Regionally, asymmetries of cortical thickness and/or surface area were found in the inferior frontal gyrus, transverse temporal gyrus, parahippocampal gyrus, and entorhinal cortex. These regions are involved in lateralized functions, including language and visuospatial processing. In addition to population-level asymmetries, variability in brain asymmetry was related to sex, age, and intracranial volume. Interestingly, we did not find significant associations between asymmetries and handedness. Finally, with two independent pedigree datasets (N = 1,443 and 1,113, respectively), we found several asymmetries showing significant, replicable heritability. The structural asymmetries identified, and their variabilities and heritability provide a reference resource for future studies on the genetic basis of brain asymmetry and altered laterality in cognitive, neurological, and psychiatric disorders.

    Additional information

    pnas.1718418115.sapp.pdf
  • Hu, C.-P., Kong, X., Wagenmakers, E.-J., Ly, A., & Peng, K. (2018). The Bayes factor and its implementation in JASP: A practical primer. Advances in Psychological Science, 26(6), 951-965. doi:10.3724/SP.J.1042.2018.00951.

    Abstract

    Statistical inference plays a critical role in modern scientific research, however, the dominant method for statistical inference in science, null hypothesis significance testing (NHST), is often misunderstood and misused, which leads to unreproducible findings. To address this issue, researchers propose to adopt the Bayes factor as an alternative to NHST. The Bayes factor is a principled Bayesian tool for model selection and hypothesis testing, and can be interpreted as the strength for both the null hypothesis H0 and the alternative hypothesis H1 based on the current data. Compared to NHST, the Bayes factor has the following advantages: it quantifies the evidence that the data provide for both the H0 and the H1, it is not “violently biased” against H0, it allows one to monitor the evidence as the data accumulate, and it does not depend on sampling plans. Importantly, the recently developed open software JASP makes the calculation of Bayes factor accessible for most researchers in psychology, as we demonstrated for the t-test. Given these advantages, adopting the Bayes factor will improve psychological researchers’ statistical inferences. Nevertheless, to make the analysis more reproducible, researchers should keep their data analysis transparent and open.
  • Kong, X., Huang, Y., Hu, S., & Liu, J. (2017). Sex-linked association between cortical scene selectivity and navigational ability. NeuroImage, 158, 397-405. doi:10.1016/j.neuroimage.2017.07.031.

    Abstract

    Spatial navigation is a crucial ability for living. Previous studies have shown that males are better at navigation than females, but little is known about the neural basis underlying the sex differences. In this study, we investigated whether cortical scene processing in three well-established scene-selective regions was sexually different, by examining sex differences in scene selectivity and its behavioral relevance to navigation. To do this, we used functional magnetic resonance imaging (fMRI) to scan the parahippocampal place area (PPA), retrosplenial complex (RSC), and occipital place area (OPA) in a large cohort of healthy young adults viewing navigationally relevant scenes (N = 202), and correlated their neural selectivity to scenes with their self-reported navigational ability. Behaviorally, we replicated the previous finding that males were better at navigation than females. Neurally, we found that the scene selectivity in the bilateral PPA, not in the RSC or OPA, was significantly higher in males than females. Such differences could not be explained by confounding factors including brain size and fMRI data quality. Importantly, males, not females, with stronger scene selectivity in the left PPA possessed better navigational ability. This brain-behavior association could not be accounted for by non-navigational abilities (i.e., intelligence and mental rotation ability). Overall, our study provides novel empirical evidence demonstrating sex differences in the brain activity, inviting further studies on sex differences in the neural network for spatial navigation.

    Additional information

    1-s2.0-S1053811917305992-mmc1.docx
  • Konopka, A., Meyer, A. S., & Forest, T. A. (2018). Planning to speak in L1 and L2. Cognitive Psychology, 102, 72-104. doi:10.1016/j.cogpsych.2017.12.003.

    Abstract

    The leading theories of sentence planning – Hierarchical Incrementality and Linear Incrementality – differ in their assumptions about the coordination of processes that map preverbal information onto language. Previous studies showed that, in native (L1) speakers, this coordination can vary with the ease of executing the message-level and sentence-level processes necessary to plan and produce an utterance. We report the first series of experiments to systematically examine how linguistic experience influences sentence planning in native (L1) speakers (i.e., speakers with life-long experience using the target language) and non-native (L2) speakers (i.e., speakers with less experience using the target language). In all experiments, speakers spontaneously generated one-sentence descriptions of simple events in Dutch (L1) and English (L2). Analyses of eye-movements across early and late time windows (pre- and post-400 ms) compared the extent of early message-level encoding and the onset of linguistic encoding. In Experiment 1, speakers were more likely to engage in extensive message-level encoding and to delay sentence-level encoding when using their L2. Experiments 2–4 selectively facilitated encoding of the preverbal message, encoding of the agent character (i.e., the first content word in active sentences), and encoding of the sentence verb (i.e., the second content word in active sentences) respectively. Experiment 2 showed that there is no delay in the onset of L2 linguistic encoding when speakers are familiar with the events. Experiments 3 and 4 showed that the delay in the onset of L2 linguistic encoding is not due to speakers delaying encoding of the agent, but due to a preference to encode information needed to select a suitable verb early in the formulation process. Overall, speakers prefer to temporally separate message-level from sentence-level encoding and to prioritize encoding of relational information when planning L2 sentences, consistent with Hierarchical Incrementality
  • Kösem, A., & Van Wassenhove, V. (2017). Distinct contributions of low and high frequency neural oscillations to speech comprehension. Language, Cognition and Neuroscience, 32(5), 536-544. doi:10.1080/23273798.2016.1238495.

    Abstract

    In the last decade, the involvement of neural oscillatory mechanisms in speech comprehension has been increasingly investigated. Current evidence suggests that low-frequency and high-frequency neural entrainment to the acoustic dynamics of speech are linked to its analysis. One crucial question is whether acoustical processing primarily modulates neural entrainment, or whether entrainment instead reflects linguistic processing. Here, we review studies investigating the effect of linguistic manipulations on neural oscillatory activity. In light of the current findings, we argue that theta (3–8 Hz) entrainment may primarily reflect the analysis of the acoustic features of speech. In contrast, recent evidence suggests that delta (1–3 Hz) and high-frequency activity (>40 Hz) are reliable indicators of perceived linguistic representations. The interdependence between low-frequency and high-frequency neural oscillations, as well as their causal role on speech comprehension, is further discussed with regard to neurophysiological models of speech processing
  • Kösem, A., Bosker, H. R., Takashima, A., Meyer, A. S., Jensen, O., & Hagoort, P. (2018). Neural entrainment determines the words we hear. Current Biology, 28, 2867-2875. doi:10.1016/j.cub.2018.07.023.

    Abstract

    Low-frequency neural entrainment to rhythmic input
    has been hypothesized as a canonical mechanism
    that shapes sensory perception in time. Neural
    entrainment is deemed particularly relevant for
    speech analysis, as it would contribute to the extraction
    of discrete linguistic elements from continuous
    acoustic signals. However, its causal influence in
    speech perception has been difficult to establish.
    Here, we provide evidence that oscillations build temporal
    predictions about the duration of speech tokens
    that affect perception. Using magnetoencephalography
    (MEG), we studied neural dynamics during
    listening to sentences that changed in speech rate.
    Weobserved neural entrainment to preceding speech
    rhythms persisting for several cycles after the change
    in rate. The sustained entrainment was associated
    with changes in the perceived duration of the last
    word’s vowel, resulting in the perception of words
    with different meanings. These findings support oscillatory
    models of speech processing, suggesting that
    neural oscillations actively shape speech perception.
  • Kotz, S. A., Ravignani, A., & Fitch, W. T. (2018). The evolution of rhythm processing. Trends in Cognitive Sciences, 22(10), 896-910. doi:10.1016/j.tics.2018.08.002.
  • Kouwenhoven, H., Van Mulken, M., & Ernestus, M. (2018). Communication strategy use by Spanish speakers of English in formal and informal speech. International Journal of Bilingualism, 22(3), 285-305. doi:10.1177/1367006916672946.

    Abstract

    Research questions:

    Are emergent bilinguals sensitive to register variation in their use of communication strategies? What strategies do LX speakers, in casu Spanish speakers of English, use as a function of situational context? What role do individual differences play?
    Methodology:

    This within-speaker study compares Spanish second-language English speakers’ communication strategy use in an informal, peer-to-peer conversation and a formal interview.
    Data and analysis:

    The 15 hours of informal and 9.5 hours of formal speech from the Nijmegen Corpus of Spanish English were coded for 19 different communication strategies.
    Findings/conclusions:

    Overall, speakers prefer self-reliant strategies, which allow them to continue communication without their interlocutor’s help. Of the self-reliant strategies, least effort strategies such as code-switching are used more often in informal speech, whereas relatively more effortful strategies (e.g. reformulations) are used more in informal speech, when the need to be unambiguously understood is felt as more important. Individual differences played a role: some speakers were more affected by a change in formality than others.
    Originality:

    Sensitivity to register variation has not yet been studied within communicative strategy use.
    Implications:

    General principles of communication govern speakers’ strategy selection, notably the protection of positive face and the least effort and cooperative principles.

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  • Kouwenhoven, H., Ernestus, M., & Van Mulken, M. (2018). Register variation by Spanish users of English. The Nijmegen Corpus of Spanish English. Corpus Linguistics and Linguistic Theory, 14(1), 35-63. doi:10.1515/cllt-2013-0054.

    Abstract

    English serves as a lingua franca in situations with varying degrees of
    formality. How formality affects non-native speech has rarely been studied. We
    investigated register variation by Spanish users of English by comparing formal
    and informal speech from the Nijmegen Corpus of Spanish English that we
    created. This corpus comprises speech from thirty-four Spanish speakers of
    English in interaction with Dutch confederates in two speech situations.
    Formality affected the amount of laughter and overlapping speech and the
    number of Spanish words. Moreover, formal speech had a more informational
    character than informal speech. We discuss how our findings relate to register
    variation in Spanish

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  • De Kovel, C. G. F., & Fisher, S. E. (2018). Molecular genetic methods. In A. M. B. De Groot, & P. Hagoort (Eds.), Research methods in psycholinguistics and the neurobiology of language: A practical guide (pp. 330-353). Hoboken: Wiley.
  • De Kovel, C. G. F., Lisgo, S., Karlebach, G., Ju, J., Cheng, G., Fisher, S. E., & Francks, C. (2017). Left-right asymmetry of maturation rates in human embryonic neural development. Biological Psychiatry, 82(3), 204-212. doi:10.1016/j.biopsych.2017.01.016.

    Abstract

    Background

    Left-right asymmetry is a fundamental organizing feature of the human brain, and neuro-psychiatric disorders such as schizophrenia sometimes involve alterations of brain asymmetry. As early as 8 weeks post conception, the majority of human fetuses move their right arms more than their left arms, but because nerve fibre tracts are still descending from the forebrain at this stage, spinal-muscular asymmetries are likely to play an important developmental role.
    Methods

    We used RNA sequencing to measure gene expression levels in the left and right spinal cords, and left and right hindbrains, of 18 post-mortem human embryos aged 4-8 weeks post conception. Genes showing embryonic lateralization were tested for an enrichment of signals in genome-wide association data for schizophrenia.
    Results

    The left side of the embryonic spinal cord was found to mature faster than the right side. Both sides transitioned from transcriptional profiles associated with cell division and proliferation at earlier stages, to neuronal differentiation and function at later stages, but the two sides were not in synchrony (p = 2.2 E-161). The hindbrain showed a left-right mirrored pattern compared to the spinal cord, consistent with the well-known crossing over of function between these two structures. Genes that showed lateralization in the embryonic spinal cord were enriched for association signals with schizophrenia (p = 4.3 E-05).
    Conclusions
    These are the earliest-stage left-right differences of human neural development ever reported. Disruption of the lateralised developmental programme may play a role in the genetic susceptibility to schizophrenia.

    Additional information

    mmc1.pdf
  • De Kovel, C. G. F., Syrbe, S., Brilstra, E. H., Verbeek, N., Kerr, B., Dubbs, H., Bayat, A., Desai, S., Naidu, S., Srivastava, S., Cagaylan, H., Yis, U., Saunders, C., Rook, M., Plugge, S., Muhle, H., Afawi, Z., Klein, K. M., Jayaraman, V., Rajagopalan, R. and 15 moreDe Kovel, C. G. F., Syrbe, S., Brilstra, E. H., Verbeek, N., Kerr, B., Dubbs, H., Bayat, A., Desai, S., Naidu, S., Srivastava, S., Cagaylan, H., Yis, U., Saunders, C., Rook, M., Plugge, S., Muhle, H., Afawi, Z., Klein, K. M., Jayaraman, V., Rajagopalan, R., Goldberg, E., Marsh, E., Kessler, S., Bergqvist, C., Conlin, L. K., Krok, B. L., Thiffault, I., Pendziwiat, M., Helbig, I., Polster, T., Borggraefe, I., Lemke, J. R., Van den Boogaardt, M. J., Moller, R. S., & Koeleman, B. P. C. (2017). Neurodevelopmental Disorders Caused by De Novo Variants in KCNB1 Genotypes and Phenotypes. JAMA Neurology, 74(10), 1228-1236. doi:10.1001/jamaneurol.2017.1714.

    Abstract

    Importance Knowing the range of symptoms seen in patients with a missense or loss-of-function variant in KCNB1 and how these symptoms correlate with the type of variant will help clinicians with diagnosis and prognosis when treating new patients. Objectives To investigate the clinical spectrum associated with KCNB1 variants and the genotype-phenotype correlations. Design, Setting, and Participants This study summarized the clinical and genetic information of patients with a presumed pathogenic variant in KCNB1.Patients were identified in research projects or during clinical testing. Information on patients from previously published articles was collected and authors contacted if feasible. All patients were seen at a clinic at one of the participating institutes because of presumed genetic disorder. They were tested in a clinical setting or included in a research project. Main Outcomes and Measures The genetic variant and its inheritance and information on the patient's symptoms and characteristics in a predefined format. All variants were identified with massive parallel sequencing and confirmed with Sanger sequencing in the patient. Absence of the variant in the parents could be confirmed with Sanger sequencing in all families except one. Results Of 26 patients (10 female, 15 male, 1 unknown; mean age at inclusion, 9.8 years; age range, 2-32 years) with developmental delay, 20 (77%) carried a missense variant in the ion channel domain of KCNB1, with a concentration of variants in region S5 to S6. Three variants that led to premature stops were located in the C-terminal and 3 in the ion channel domain. Twenty-one of 25 patients (84%) had seizures, with 9 patients (36%) starting with epileptic spasms between 3 and 18 months of age. All patients had developmental delay, with 17 (65%) experiencing severe developmental delay; 14 (82%) with severe delay had behavioral problems. The developmental delay was milder in 4 of 6 patients with stop variants and in a patient with a variant in the S2 transmembrane element rather than the S4 to S6 region. Conclusions and Relevance De novo KCNB1 missense variants in the ion channel domain and loss-of-function variants in this domain and the C-terminal likely cause neurodevelopmental disorders with or without seizures. Patients with presumed pathogenic variants in KCNB1 have a variable phenotype. However, the type and position of the variants in the protein are (imperfectly) correlated with the severity of the disorder.
  • De Kovel, C. G. F., Lisgo, S. N., Fisher, S. E., & Francks, C. (2018). Subtle left-right asymmetry of gene expression profiles in embryonic and foetal human brains. Scientific Reports, 8: 12606. doi:10.1038/s41598-018-29496-2.

    Abstract

    Left-right laterality is an important aspect of human –and in fact all vertebrate– brain organization for which the genetic basis is poorly understood. Using RNA sequencing data we contrasted gene expression in left- and right-sided samples from several structures of the anterior central nervous systems of post mortem human embryos and foetuses. While few individual genes stood out as significantly lateralized, most structures showed evidence of laterality of their overall transcriptomic profiles. These left-right differences showed overlap with age-dependent changes in expression, indicating lateralized maturation rates, but not consistently in left-right orientation over all structures. Brain asymmetry may therefore originate in multiple locations, or if there is a single origin, it is earlier than 5 weeks post conception, with structure-specific lateralized processes already underway by this age. This pattern is broadly consistent with the weak correlations reported between various aspects of adult brain laterality, such as language dominance and handedness.
  • De Kovel, C. G. F., Lisgo, S. N., & Francks, C. (2018). Transcriptomic analysis of left-right differences in human embryonic forebrain and midbrain. Scientific Data, 5: 180164. doi:10.1038/sdata.2018.164.

    Abstract

    Left-right asymmetry is subtle but pervasive in the human central nervous system. This asymmetry is initiated early during development, but its mechanisms are poorly known. Forebrains and midbrains were dissected from six human embryos at Carnegie stages 15 or 16, one of which was female. The structures were divided into left and right sides, and RNA was isolated. RNA was sequenced with 100 base-pair paired ends using Illumina Hiseq 4000. After quality control, five paired brain sides were available for midbrain and forebrain. A paired analysis between left- and right sides of a given brain structure across the embryos identified left-right differences. The dataset, consisting of Fastq files and a read count table, can be further used to study early development of the human brain
  • Kuerbitz, J., Arnett, M., Ehrman, S., Williams, M. T., Voorhees, C. V., Fisher, S. E., Garratt, A. N., Muglia, L. J., Waclaw, R. R., & Campbell, K. (2018). Loss of intercalated cells (ITCs) in the mouse amygdala of Tshz1 mutants correlates with fear, depression and social interaction phenotypes. The Journal of Neuroscience, 38, 1160-1177. doi:10.1523/JNEUROSCI.1412-17.2017.

    Abstract

    The intercalated cells (ITCs) of the amygdala have been shown to be critical regulatory components of amygdalar circuits, which control appropriate fear responses. Despite this, the molecular processes guiding ITC development remain poorly understood. Here we establish the zinc finger transcription factor Tshz1 as a marker of ITCs during their migration from the dorsal lateral ganglionic eminence through maturity. Using germline and conditional knock-out (cKO) mouse models, we show that Tshz1 is required for the proper migration and differentiation of ITCs. In the absence of Tshz1, migrating ITC precursors fail to settle in their stereotypical locations encapsulating the lateral amygdala and BLA. Furthermore, they display reductions in the ITC marker Foxp2 and ectopic persistence of the dorsal lateral ganglionic eminence marker Sp8. Tshz1 mutant ITCs show increased cell death at postnatal time points, leading to a dramatic reduction by 3 weeks of age. In line with this, Foxp2-null mutants also show a loss of ITCs at postnatal time points, suggesting that Foxp2 may function downstream of Tshz1 in the maintenance of ITCs. Behavioral analysis of male Tshz1 cKOs revealed defects in fear extinction as well as an increase in floating during the forced swim test, indicative of a depression-like phenotype. Moreover, Tshz1 cKOs display significantly impaired social interaction (i.e., increased passivity) regardless of partner genetics. Together, these results suggest that Tshz1 plays a critical role in the development of ITCs and that fear, depression-like and social behavioral deficits arise in their absence. SIGNIFICANCE STATEMENT We show here that the zinc finger transcription factor Tshz1 is expressed during development of the intercalated cells (ITCs) within the mouse amygdala. These neurons have previously been shown to play a crucial role in fear extinction. Tshz1 mouse mutants exhibit severely reduced numbers of ITCs as a result of abnormal migration, differentiation, and survival of these neurons. Furthermore, the loss of ITCs in mouse Tshz1 mutants correlates well with defects in fear extinction as well as the appearance of depression-like and abnormal social interaction behaviors reminiscent of depressive disorders observed in human patients with distal 18q deletions, including the Tshz1 locus.
  • Kuiper, K., Bimesl, N., Kempen, G., & Ogino, M. (2017). Initial vs. non-initial placement of agent constructions in spoken clauses: A corpus-based study of language production under time pressure. Language Sciences, 64, 16-33. doi:10.1016/j.langsci.2017.06.001.

    Abstract

    In this exploratory study we test the hypothesis that the retrieval from memory of proper noun Agents (PNAs) under processing pressure causes a greater proportion of such semantic arguments to be placed to the right of the initial position in a clause than would be the case if such retrieval from memory were not necessary. This effect is manifest in sports commentary. Processing pressure on sports commentators is modulated by the speed at which the sport is played and reported. Non-initial placement is also facilitated by formulae which have slots in non-initial position. It follows that the non-initial placement of PNAs is not always semantically or pragmatically motivated. This finding therefore runs counter to a strong form of the functionalist hypothesis that syntactic choices available in the systemic structure of the syntax of a language offer solely semantic or pragmatic choices. It is an open question in a weak functionalist account of language and language use how processing and communicative functions interact in general.
  • Kunert, R., & Jongman, S. R. (2017). Entrainment to an auditory signal: Is attention involved? Journal of Experimental Psychology: General, 146(1), 77-88. doi:10.1037/xge0000246.

    Abstract

    Many natural auditory signals, including music and language, change periodically. The effect of such auditory rhythms on the brain is unclear however. One widely held view, dynamic attending theory, proposes that the attentional system entrains to the rhythm and increases attention at moments of rhythmic salience. In support, 2 experiments reported here show reduced response times to visual letter strings shown at auditory rhythm peaks, compared with rhythm troughs. However, we argue that an account invoking the entrainment of general attention should further predict rhythm entrainment to also influence memory for visual stimuli. In 2 pseudoword memory experiments we find evidence against this prediction. Whether a pseudoword is shown during an auditory rhythm peak or not is irrelevant for its later recognition memory in silence. Other attention manipulations, dividing attention and focusing attention, did result in a memory effect. This raises doubts about the suggested attentional nature of rhythm entrainment. We interpret our findings as support for auditory rhythm perception being based on auditory-motor entrainment, not general attention entrainment.
  • Kunert, R. (2017). Music and language comprehension in the brain. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Kung, C. (2018). Speech comprehension in a tone language: The role of lexical tone, context, and intonation in Cantonese-Chinese. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Lakens, D., Adolfi, F. G., Albers, C. J., Anvari, F., Apps, M. A. J., Argamon, S. E., Baguley, T., Becker, R. B., Benning, S. D., Bradford, D. E., Buchanan, E. M., Caldwell, A. R., Van Calster, B., Carlsson, R., Chen, S.-C., Chung, B., Colling, L. J., Collins, G. S., Crook, Z., Cross, E. S. and 68 moreLakens, D., Adolfi, F. G., Albers, C. J., Anvari, F., Apps, M. A. J., Argamon, S. E., Baguley, T., Becker, R. B., Benning, S. D., Bradford, D. E., Buchanan, E. M., Caldwell, A. R., Van Calster, B., Carlsson, R., Chen, S.-C., Chung, B., Colling, L. J., Collins, G. S., Crook, Z., Cross, E. S., Daniels, S., Danielsson, H., DeBruine, L., Dunleavy, D. J., Earp, B. D., Feist, M. I., Ferrelle, J. D., Field, J. G., Fox, N. W., Friesen, A., Gomes, C., Gonzalez-Marquez, M., Grange, J. A., Grieve, A. P., Guggenberger, R., Grist, J., Van Harmelen, A.-L., Hasselman, F., Hochard, K. D., Hoffarth, M. R., Holmes, N. P., Ingre, M., Isager, P. M., Isotalus, H. K., Johansson, C., Juszczyk, K., Kenny, D. A., Khalil, A. A., Konat, B., Lao, J., Larsen, E. G., Lodder, G. M. A., Lukavský, J., Madan, C. R., Manheim, D., Martin, S. R., Martin, A. E., Mayo, D. G., McCarthy, R. J., McConway, K., McFarland, C., Nio, A. Q. X., Nilsonne, G., De Oliveira, C. L., De Xivry, J.-J.-O., Parsons, S., Pfuhl, G., Quinn, K. A., Sakon, J. J., Saribay, S. A., Schneider, I. K., Selvaraju, M., Sjoerds, Z., Smith, S. G., Smits, T., Spies, J. R., Sreekumar, V., Steltenpohl, C. N., Stenhouse, N., Świątkowski, W., Vadillo, M. A., Van Assen, M. A. L. M., Williams, M. N., Williams, S. E., Williams, D. R., Yarkoni, T., Ziano, I., & Zwaan, R. A. (2018). Justify your alpha. Nature Human Behaviour, 2, 168-171. doi:10.1038/s41562-018-0311-x.

    Abstract

    In response to recommendations to redefine statistical significance to P ≤ 0.005, we propose that researchers should transparently report and justify all choices they make when designing a study, including the alpha level.
  • Lam, N. H. L. (2017). Comprehending comprehension: Insights from neuronal oscillations on the neuronal basis of language. PhD Thesis, Radboud University Nijmegen, Nijmegen.
  • Lam, K. J. Y., Bastiaansen, M. C. M., Dijkstra, T., & Rueschemeyer, S. A. (2017). Making sense: motor activation and action plausibility during sentence processing. Language, Cognition and Neuroscience, 32(5), 590-600. doi:10.1080/23273798.2016.1164323.

    Abstract

    The current electroencephalography study investigated the relationship between the motor and (language) comprehension systems by simultaneously measuring mu and N400 effects. Specifically, we examined whether the pattern of motor activation elicited by verbs depends on the larger sentential context. A robust N400 congruence effect confirmed the contextual manipulation of action plausibility, a form of semantic congruency. Importantly, this study showed that: (1) Action verbs elicited more mu power decrease than non-action verbs when sentences described plausible actions. Action verbs thus elicited more motor activation than non-action verbs. (2) In contrast, when sentences described implausible actions, mu activity was present but the difference between the verb types was not observed. The increased processing associated with a larger N400 thus coincided with mu activity in sentences describing implausible actions. Altogether, context-dependent motor activation appears to play a functional role in deriving context-sensitive meaning
  • Lam, N. H. L., Hulten, A., Hagoort, P., & Schoffelen, J.-M. (2018). Robust neuronal oscillatory entrainment to speech displays individual variation in lateralisation. Language, Cognition and Neuroscience, 33(8), 943-954. doi:10.1080/23273798.2018.1437456.

    Abstract

    Neural oscillations may be instrumental for the tracking and segmentation of continuous speech. Earlier work has suggested that delta, theta and gamma oscillations entrain to the speech rhythm. We used magnetoencephalography and a large sample of 102 participants to investigate oscillatory entrainment to speech, and observed robust entrainment of delta and theta activity, and weak group-level gamma entrainment. We show that the peak frequency and the hemispheric lateralisation of the entrainment are subject to considerable individual variability. The first finding may support the involvement of intrinsic oscillations in entrainment, and the second finding suggests that there is no systematic default right-hemispheric bias for processing acoustic signals on a slow time scale. Although low frequency entrainment to speech is a robust phenomenon, the characteristics of entrainment vary across individuals, and this variation is important for understanding the underlying neural mechanisms of entrainment, as well as its functional significance.
  • Lattenkamp, E. Z., Kaiser, S., Kaucic, R., Großmann, M., Koselj, K., & Goerlitz, H. R. (2018). Environmental acoustic cues guide the biosonar attention of a highly specialised echolocator. Journal of Experimental Biology, 221(8): jeb165696. doi:10.1242/jeb.165696.

    Abstract

    Sensory systems experience a trade-off between maximizing the
    detail and amount of sampled information. Thistrade-off is particularly
    pronounced in sensorysystemsthat are highlyspecialised fora single
    task and thus experience limitations in other tasks. We hypothesised
    that combining sensory input from multiple streams of information
    may resolve this trade-off and improve detection and sensing
    reliability. Specifically, we predicted that perceptive limitations
    experienced by animals reliant on specialised active echolocation
    can be compensated for by the phylogenetically older and less
    specialised process of passive hearing. We tested this hypothesis in
    greater horseshoe bats, which possess morphological and neural
    specialisations allowing them to identify fluttering prey in dense
    vegetation using echolocation only. At the same time, their
    echolocation system is both spatially and temporally severely
    limited. Here, we show that greater horseshoe bats employ passive
    hearing to initially detect and localise prey-generated and other
    environmental sounds, and then raise vocalisation level and
    concentrate the scanning movements of their sonar beam on the
    sound source for further investigation with echolocation. These
    specialised echolocators thus supplement echo-acoustic information
    with environmental acoustic cues, enlarging perceived space beyond
    their biosonar range. Contrary to our predictions, we did not find
    consistent preferences for prey-related acoustic stimuli, indicating the
    use of passive acoustic cues also for detection of non-prey objects.
    Our findings suggest that even specialised echolocators exploit a
    wide range of environmental information, and that phylogenetically
    older sensory systems can support the evolution of sensory
    specialisations by compensating for their limitations.
  • Lattenkamp, E. Z., Vernes, S. C., & Wiegrebe, L. (2018). Mammalian models for the study of vocal learning: A new paradigm in bats. In C. Cuskley, M. Flaherty, H. Little, L. McCrohon, A. Ravignani, & T. Verhoef (Eds.), Proceedings of the 12th International Conference on the Evolution of Language (EVOLANG XII) (pp. 235-237). Toruń, Poland: NCU Press. doi:10.12775/3991-1.056.
  • Lattenkamp, E. Z., & Vernes, S. C. (2018). Vocal learning: A language-relevant trait in need of a broad cross-species approach. Current Opinion in Behavioral Sciences, 21, 209-215. doi:10.1016/j.cobeha.2018.04.007.

    Abstract

    Although humans are unmatched in their capacity to produce
    speech and learn language, comparative approaches in diverse
    animalmodelsareabletoshedlightonthebiologicalunderpinnings
    of language-relevant traits. In the study of vocal learning, a trait
    crucial for spoken language, passerine birds have been the
    dominant models, driving invaluable progress in understanding the
    neurobiology and genetics of vocal learning despite being only
    distantly related to humans. To date, there is sparse evidence that
    our closest relatives, nonhuman primates have the capability to
    learn new vocalisations. However, a number of other mammals
    have shown the capacity for vocal learning, such as some
    cetaceans, pinnipeds, elephants, and bats, and we anticipate that
    with further study more species will gain membership to this
    (currently) select club. A broad, cross-species comparison of vocal
    learning, coupled with careful consideration of the components
    underlying this trait, is crucial to determine how human speech and
    spoken language is biologically encoded and how it evolved. We
    emphasise the need to draw on the pool of promising species that
    havethusfarbeenunderstudiedorneglected.Thisisbynomeansa
    call for fewer studies in songbirds, or an unfocused treasure-hunt,
    but rather an appeal for structured comparisons across a range of
    species, considering phylogenetic relationships, ecological and
    morphological constrains, developmental and social factors, and
    neurogenetic underpinnings. Herein, we promote a comparative
    approachhighlightingtheimportanceofstudyingvocallearningina
    broad range of model species, and describe a common framework
    for targeted cross-taxon studies to shed light on the biology and
    evolution of vocal learning.
  • Lattenkamp, E. Z., Vernes, S. C., & Wiegrebe, L. (2018). Volitional control of social vocalisations and vocal usage learning in bats. Journal of Experimental Biology, 221(14): jeb.180729. doi:10.1242/jeb.180729.

    Abstract

    Bats are gregarious, highly vocal animals that possess a broad repertoire of social vocalisations. For in-depth studies of their vocal behaviours, including vocal flexibility and vocal learning, it is necessary to gather repeatable evidence from controlled laboratory experiments on isolated individuals. However, such studies are rare for one simple reason: eliciting social calls in isolation and under operant control is challenging and has rarely been achieved. To overcome this limitation, we designed an automated setup that allows conditioning of social vocalisations in a new context, and tracks spectro-temporal changes in the recorded calls over time. Using this setup, we were able to reliably evoke social calls from temporarily isolated lesser spear-nosed bats (Phyllostomus discolor). When we adjusted the call criteria that could result in food reward, bats responded by adjusting temporal and spectral call parameters. This was achieved without the help of an auditory template or social context to direct the bats. Our results demonstrate vocal flexibility and vocal usage learning in bats. Our setup provides a new paradigm that allows the controlled study of the production and learning of social vocalisations in isolated bats, overcoming limitations that have, until now, prevented in-depth studies of these behaviours.

    Additional information

    JEB180729supp.pdf
  • Lauscher, A., Eckert, K., Galke, L., Scherp, A., Rizvi, S. T. R., Ahmed, S., Dengel, A., Zumstein, P., & Klein, A. (2018). Linked open citation database: Enabling libraries to contribute to an open and interconnected citation graph. In J. Chen, M. A. Gonçalves, J. M. Allen, E. A. Fox, M.-Y. Kan, & V. Petras (Eds.), JCDL '18: Proceedings of the 18th ACM/IEEE on Joint Conference on Digital Libraries (pp. 109-118). New York: ACM. doi:10.1145/3197026.3197050.

    Abstract

    Citations play a crucial role in the scientific discourse, in information retrieval, and in bibliometrics. Many initiatives are currently promoting the idea of having free and open citation data. Creation of citation data, however, is not part of the cataloging workflow in libraries nowadays.
    In this paper, we present our project Linked Open Citation Database, in which we design distributed processes and a system infrastructure based on linked data technology. The goal is to show that efficiently cataloging citations in libraries using a semi-automatic approach is possible. We specifically describe the current state of the workflow and its implementation. We show that we could significantly improve the automatic reference extraction that is crucial for the subsequent data curation. We further give insights on the curation and linking process and provide evaluation results that not only direct the further development of the project, but also allow us to discuss its overall feasibility.
  • Lee, R., Chambers, C. G., Huettig, F., & Ganea, P. A. (2017). Children’s semantic and world knowledge overrides fictional information during anticipatory linguistic processing. In G. Gunzelmann, A. Howes, T. Tenbrink, & E. Davelaar (Eds.), Proceedings of the 39th Annual Meeting of the Cognitive Science Society (CogSci 2017) (pp. 730-735). Austin, TX: Cognitive Science Society.

    Abstract

    Using real-time eye-movement measures, we asked how a fantastical discourse context competes with stored representations of semantic and world knowledge to influence children's and adults' moment-by-moment interpretation of a story. Seven-year- olds were less effective at bypassing stored semantic and world knowledge during real-time interpretation than adults. Nevertheless, an effect of discourse context on comprehension was still apparent.
  • Lee, J. J., Wedow, R., Okbay, A., Kong, E., Maghzian, O., Zacher, M., Nguyen-Viet, T. A., Bowers, P., Sidorenko, J., Linnér, R. K., Fontana, M. A., Kundu, T., Lee, C., Li, H., Li, R., Royer, R., Timshel, P. N., Walters, R. K., Willoughby, E. A., Yengo, L. and 57 moreLee, J. J., Wedow, R., Okbay, A., Kong, E., Maghzian, O., Zacher, M., Nguyen-Viet, T. A., Bowers, P., Sidorenko, J., Linnér, R. K., Fontana, M. A., Kundu, T., Lee, C., Li, H., Li, R., Royer, R., Timshel, P. N., Walters, R. K., Willoughby, E. A., Yengo, L., 23andMe Research Team, COGENT (Cognitive Genomics Consortium), Social Science Genetic Association Consortium, Alver, M., Bao, Y., Clark, D. W., Day, F. R., Furlotte, N. A., Joshi, P. K., Kemper, K. E., Kleinman, A., Langenberg, C., Mägi, R., Trampush, J. W., Verma, S. S., Wu, Y., Lam, M., Zhao, J. H., Zheng, Z., Boardman, J. D., Campbell, H., Freese, J., Harris, K. M., Hayward, C., Herd, P., Kumari, M., Lencz, T., Luan, J., Malhotra, A. K., Metspalu, A., Milani, L., Ong, K. K., Perry, J. R. B., Porteous, D. J., Ritchie, M. D., Smart, M. C., Smith, B. H., Tung, J. Y., Wareham, N. J., Wilson, J. F., Beauchamp, J. P., Conley, D. C., Esko, T., Lehrer, S. F., Magnusson, P. K. E., Oskarsson, S., Pers, T. H., Robinson, M. R., Thom, K., Watson, C., Chabris, C. F., Meyer, M. N., Laibson, D. I., Yang, J., Johannesson, M., Koellinger, P. D., Turley, P., Visscher, P. M., Benjamin, D. J., & Cesarini, D. (2018). Gene discovery and polygenic prediction from a genome-wide association study of educational attainment in 1.1 million individuals. Nature Genetics, 50(8), 1112-1121. doi:10.1038/s41588-018-0147-3.

    Abstract

    Here we conducted a large-scale genetic association analysis of educational attainment in a sample of approximately 1.1 million individuals and identify 1,271 independent genome-wide-significant SNPs. For the SNPs taken together, we found evidence of heterogeneous effects across environments. The SNPs implicate genes involved in brain-development processes and neuron-to-neuron communication. In a separate analysis of the X chromosome, we identify 10 independent genome-wide-significant SNPs and estimate a SNP heritability of around 0.3% in both men and women, consistent with partial dosage compensation. A joint (multi-phenotype) analysis of educational attainment and three related cognitive phenotypes generates polygenic scores that explain 11–13% of the variance in educational attainment and 7–10% of the variance in cognitive performance. This prediction accuracy substantially increases the utility of polygenic scores as tools in research.
  • Lefever, E., Hendrickx, I., Croijmans, I., Van den Bosch, A., & Majid, A. (2018). Discovering the language of wine reviews: A text mining account. In N. Calzolari, K. Choukri, C. Cieri, T. Declerck, S. Goggi, K. Hasida, H. Isahara, B. Maegaard, J. Mariani, H. Mazo, A. Moreno, J. Odijk, S. Piperidis, & T. Tokunaga (Eds.), Proceedings of the Eleventh International Conference on Language Resources and Evaluation (LREC 2018) (pp. 3297-3302). Paris: LREC.

    Abstract

    It is widely held that smells and flavors are impossible to put into words. In this paper we test this claim by seeking predictive patterns in wine reviews, which ostensibly aim to provide guides to perceptual content. Wine reviews have previously been critiqued as random and meaningless. We collected an English corpus of wine reviews with their structured metadata, and applied machine learning techniques to automatically predict the wine's color, grape variety, and country of origin. To train the three supervised classifiers, three different information sources were incorporated: lexical bag-of-words features, domain-specific terminology features, and semantic word embedding features. In addition, using regression analysis we investigated basic review properties, i.e., review length, average word length, and their relationship to the scalar values of price and review score. Our results show that wine experts do share a common vocabulary to describe wines and they use this in a consistent way, which makes it possible to automatically predict wine characteristics based on the review text alone. This means that odors and flavors may be more expressible in language than typically acknowledged.
  • Lemhöfer, K., Huestegge, L., & Mulder, K. (2018). Another cup of TEE? The processing of second language near-cognates in first language reading. Language, Cognition and Neuroscience, 33(8), 968-991. doi:10.1080/23273798.2018.1433863.

    Abstract

    A still unresolved issue is in how far native language (L1) processing in bilinguals is influenced by the second language (L2). We investigated this in two word recognition experiments in L1, using homophonic near-cognates that are spelled in L2. In a German lexical decision task (Experiment 1), German-Dutch bilinguals had more difficulties to reject these Dutch-spelled near-cognates than other misspellings, while this was not the case for non-Dutch speaking Germans. In Experiment 2, the same materials were embedded in German sentences. Analyses of eye movements during reading showed that only non-Dutch speaking Germans, but not Dutch-speaking participants were slowed down by the Dutch cognate misspellings. Additionally, in both experiments, bilinguals with larger vocabulary sizes in Dutch tended to show larger near-cognate effects. Thus, Dutch word knowledge influenced word recognition in L1 German in both task contexts, suggesting that L1 word recognition in bilinguals is non-selective with respect to L2.
  • Lev-Ari, S., & Shao, Z. (2017). How social network heterogeneity facilitates lexical access and lexical prediction. Memory & Cognition, 45(3), 528-538. doi:10.3758/s13421-016-0675-y.

    Abstract

    People learn language from their social environment. As individuals differ in their social networks, they might be exposed to input with different lexical distributions, and these might influence their linguistic representations and lexical choices. In this article we test the relation between linguistic performance and 3 social network properties that should influence input variability, namely, network size, network heterogeneity, and network density. In particular, we examine how these social network properties influence lexical prediction, lexical access, and lexical use. To do so, in Study 1, participants predicted how people of different ages would name pictures, and in Study 2 participants named the pictures themselves. In both studies, we examined how participants’ social network properties related to their performance. In Study 3, we ran simulations on norms we collected to see how age variability in one’s network influences the distribution of different names in the input. In all studies, network age heterogeneity influenced performance leading to better prediction, faster response times for difficult-to-name items, and less entropy in input distribution. These results suggest that individual differences in social network properties can influence linguistic behavior. Specifically, they show that having a more heterogeneous network is associated with better performance. These results also show that the same factors influence lexical prediction and lexical production, suggesting the two might be related.
  • Lev-Ari, S., & Peperkamp, S. (2017). Language for $200: Success in the environment influences grammatical alignment. Journal of Language Evolution, 2(2), 177-187. doi:10.1093/jole/lzw012.

    Abstract

    Speakers constantly learn language from the environment by sampling their linguistic input and adjusting their representations accordingly. Logically, people should attend more to the environment and adjust their behavior in accordance with it more the lower their success in the environment is. We test whether the learning of linguistic input follows this general principle in two studies: a corpus analysis of a TV game show, Jeopardy, and a laboratory task modeled after Go Fish. We show that lower (non-linguistic) success in the task modulates learning of and reliance on linguistic patterns in the environment. In Study 1, we find that poorer performance increases conformity with linguistic norms, as reflected by increased preference for frequent grammatical structures. In Study 2, which consists of a more interactive setting, poorer performance increases learning from the immediate social environment, as reflected by greater repetition of others’ grammatical structures. We propose that these results have implications for models of language production and language learning and for the propagation of language change. In particular, they suggest that linguistic changes might spread more quickly in times of crisis, or when the gap between more and less successful people is larger. The results might also suggest that innovations stem from successful individuals while their propagation would depend on relatively less successful individuals. We provide a few historical examples that are in line with the first suggested implication, namely, that the spread of linguistic changes is accelerated during difficult times, such as war time and an economic downturn
  • Lev-Ari, S., van Heugten, M., & Peperkamp, S. (2017). Relative difficulty of understanding foreign accents as a marker of proficiency. Cognitive Science, 41(4), 1106-1118. doi:10.1111/cogs.12394.

    Abstract

    Foreign-accented speech is generally harder to understand than native-accented speech. This difficulty is reduced for non-native listeners who share their first language with the non-native speaker. It is currently unclear, however, how non-native listeners deal with foreign-accented speech produced by speakers of a different language. We show that the process of (second) language acquisition is associated with an increase in the relative difficulty of processing foreign-accented speech. Therefore, experiencing greater relative difficulty with foreign-accented speech compared with native speech is a marker of language proficiency. These results contribute to our understanding of how phonological categories are acquired during second language learning.
  • Lev-Ari, S. (2017). Talking to fewer people leads to having more malleable linguistic representations. PLoS One, 12(8): e0183593. doi:10.1371/journal.pone.0183593.

    Abstract

    We learn language from our social environment. In general, the more sources we have, the less informative each of them is, and the less weight we should assign it. If this is the case, people who interact with fewer others should be more susceptible to the influence of each of their interlocutors. This paper tests whether indeed people who interact with fewer other people have more malleable phonological representations. Using a perceptual learning paradigm, this paper shows that individuals who regularly interact with fewer others are more likely to change their boundary between /d/ and /t/ following exposure to an atypical speaker. It further shows that the effect of number of interlocutors is not due to differences in ability to learn the speaker’s speech patterns, but specific to likelihood of generalizing the learned pattern. These results have implications for both language learning and language change, as they suggest that individuals with smaller social networks might play an important role in propagating linguistic changes.

    Additional information

    5343619.zip
  • Lev-Ari, S. (2018). Social network size can influence linguistic malleability and the propagation of linguistic change. Cognition, 176, 31-39. doi:10.1016/j.cognition.2018.03.003.

    Abstract

    We learn language from our social environment, but the more sources we have, the less informative each source is, and therefore, the less weight we ascribe its input. According to this principle, people with larger social networks should give less weight to new incoming information, and should therefore be less susceptible to the influence of new speakers. This paper tests this prediction, and shows that speakers with smaller social networks indeed have more malleable linguistic representations. In particular, they are more likely to adjust their lexical boundary following exposure to a new speaker. Experiment 2 uses computational simulations to test whether this greater malleability could lead people with smaller social networks to be important for the propagation of linguistic change despite the fact that they interact with fewer people. The results indicate that when innovators were connected with people with smaller rather than larger social networks, the population exhibited greater and faster diffusion. Together these experiments show that the properties of people’s social networks can influence individuals’ learning and use as well as linguistic phenomena at the community level.
  • Lev-Ari, S. (2018). The influence of social network size on speech perception. Quarterly Journal of Experimental Psychology, 71(10), 2249-2260. doi:10.1177/1747021817739865.

    Abstract

    Infants and adults learn new phonological varieties better when exposed to multiple rather than a single speaker. This
    article tests whether having a larger social network similarly facilitates phonological performance. Experiment 1 shows
    that people with larger social networks are better at vowel perception in noise, indicating that the benefit of laboratory
    exposure to multiple speakers extends to real life experience and to adults tested in their native language. Furthermore,
    the experiment shows that this association is not due to differences in amount of input or to cognitive differences
    between people with different social network sizes. Follow-up computational simulations reveal that the benefit of
    larger social networks is mostly due to increased input variability. Additionally, the simulations show that the boost
    that larger social networks provide is independent of the amount of input received but is larger if the population is
    more heterogeneous. Finally, a comparison of “adult” and “child” simulations reconciles previous conflicting findings
    by suggesting that input variability along the relevant dimension might be less useful at the earliest stages of learning.
    Together, this article shows when and how the size of our social network influences our speech perception. It thus
    shows how aspects of our lifestyle can influence our linguistic performance.

    Additional information

    QJE-STD_17-073.R4-Table_A1.docx
  • Lev-Ari, S., Ho, E., & Keysar, B. (2018). The unforeseen consequences of interacting with non-native speakers. Topics in Cognitive Science, 10, 835-849. doi:10.1111/tops.12325.

    Abstract

    Sociolinguistic research shows that listeners' expectations of speakers influence their interpretation of the speech, yet this is often ignored in cognitive models of language comprehension. Here, we focus on the case of interactions between native and non-native speakers. Previous literature shows that listeners process the language of non-native speakers in less detail, because they expect them to have lower linguistic competence. We show that processing the language of non-native speakers increases lexical competition and access in general, not only of the non-native speaker's speech, and that this leads to poorer memory of one's own speech during the interaction. We further find that the degree to which people adjust their processing to non-native speakers is related to the degree to which they adjust their speech to them. We discuss implications for cognitive models of language processing and sociolinguistic research on attitudes.
  • Levelt, W. J. M. (1982). Cognitive styles in the use of spatial direction terms. In R. Jarvella, & W. Klein (Eds.), Speech, place, and action: Studies in deixis and related topics (pp. 251-268). Chichester: Wiley.
  • Levelt, W. J. M. (1976). Formal grammars and the natural language user: A review. In A. Marzollo (Ed.), Topics in artificial intelligence (pp. 226-290). Vienna: Springer.
  • Levelt, W. J. M. (1973). Formele grammatica's in linguistiek en taalpsychologie (Vols. I-III). Deventer: Van Loghem Slaterus.
  • Levelt, W. J. M. (1962). Motion breaking and the perception of causality. In A. Michotte (Ed.), Causalité, permanence et réalité phénoménales: Etudes de psychologie expérimentale (pp. 244-258). Louvain: Publications Universitaires.
  • Levelt, W. J. M., & Plomp, R. (1962). Musical consonance and critical bandwidth. In Proceedings of the 4th International Congress Acoustics (pp. 55-55).
  • Levelt, W. J. M. (1982). Het lineariseringsprobleem van de spreker. Tijdschrift voor Taal- en Tekstwetenschap (TTT), 2(1), 1-15.
  • Levelt, W. J. M. (1982). Linearization in describing spatial networks. In S. Peters, & E. Saarinen (Eds.), Processes, beliefs, and questions (pp. 199-220). Dordrecht - Holland: D. Reidel.

    Abstract

    The topic of this paper is the way in which speakers order information in discourse. I will refer to this issue with the term "linearization", and will begin with two types of general remarks. The first one concerns the scope and relevance of the problem with reference to some existing literature. The second set of general remarks will be about the place of linearization in a theory of the speaker. The following, and main part of this paper, will be a summary report of research of linearization in a limited, but well-defined domain of discourse, namely the description of spatial networks.
  • Levelt, W. J. M. (2018). Is language natural to man? Some historical considerations. Current Opinion in Behavioral Sciences, 21, 127-131. doi:10.1016/j.cobeha.2018.04.003.

    Abstract

    Since the Enlightenment period, natural theories of speech and language evolution have florished in the language sciences. Four ever returning core issues are highlighted in this paper: Firstly, Is language natural to man or just an invention? Secondly, Is language a specific human ability (a ‘language instinct’) or does it arise from general cognitive capacities we share with other animals? Thirdly, Has the evolution of language been a gradual process or did it rather suddenly arise, due to some ‘evolutionary twist’? Lastly, Is the child's language acquisition an appropriate model for language evolution?
  • Levelt, W. J. M. (1973). Recente ontwikkelingen in de taalpsychologie. Forum der Letteren, 14(4), 235-254.

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